Cladistic analysis and synopsis of Chloropepla Stål (Hemiptera: Heteroptera: Pentatomidae) with the description of three new species
Author
Grevea, Caroline
Universidade Federal do Rio Grande do Sul, Departamento de Zoologia, Av. Bento Gonçalves 9.500, 91501 - 970 Porto Alegre, Rio Grande do Sul, Brazil
carolinegreve@yahoo.com.br
Author
Schwertnerb, Cristiano F.
Laboratório de Ecologia e Sistemática, Departamento de Ciências Biológicas, Programa de Pós-Graduação em Ecologia e Evolução, Universidade de São Paulo, Rua Prof. Artur Riedel 275, 09972 – 270 Diadema-SP, Brazil
Author
Grazia, Jocelia
Universidade Federal do Rio Grande do Sul, Departamento de Zoologia, Av. Bento Gonçalves 9.500, 91501 - 970 Porto Alegre, Rio Grande do Sul, Brazil
text
Insect Systematics & Evolution
2013
2013-04-15
44
1
43
journal article
10.1163/1876312X-04401002
e2ed26bf-60f8-4dca-b769-b6029e9e00fa
3653979
Key to
Chloropepla
species
1 Humeral angles produced into spines (
Fig. 1C
) ................................................. 2
- Humeral angles acute but not produced into spines (
Fig. 1E
)............................ 4
2 Humeral angles outlined in black (
Fig. 1D
)....................................................... 3
- Humeral angles not outlined in black (
Fig. 1C, E
) .......................................... 12
3 Lateral margin of each jugum uniformly convex towards apex (
Fig. 1A
), bordered in black; dorsum of each tibia outlined in black ................
C. aurea
(
Pirán, 1963
)
- Lateral margins of juga sinuous (
Fig. 1B
), not bordered in black; dorsum of tibia not outlined in black ..............................................
C. pirani
Grazia-Vieira, 1971
4 Apices of femora with conspicuous spine (
Fig. 2G
) ........................................... 5
- Apices of femora with reduced spine (
Fig. 2F
) ................................................... 6
5 Hypandrium with 1+ 1 broad expansions flanking segment X, each expansion bearing an elongated outgrowth on dorsal surface (
Fig. 11A
); posterior margin of gonocoxites 9 straight (
Fig. 12A
) ......................................
Chloropepla rideri
sp.n
.
- Hypandrium with 1 +1 narrow expansions flanking segment X, without outgrowth on dorsal surface (
Fig. 8A
); posterior margin of gonocoxites 9 convex (
Fig. 8B
)..........................................
C. paveli
Grazia, Schwertner & Greve, 2008
6 Medial excavation in ventral rim of pygophore “V”-shaped (
Fig. 4F
); marginal process of dorsal rim of pygophore triangular (
Fig. 4A, B
); gonocoxites 9 strongly convex, with lateral margins nearly parallel (
Fig. 16A
) ....................................... 7
- Medial excavation in ventral rim of pygophore “U”-shaped (
Fig. 4G
); marginal process of dorsal rim of pygophore digitiform (
Fig. 4C-E
), gonocoxites 9 slightly convex, with lateral margins convergent ............................................................ 8
7 Apex of hypandrium rounded, without process (
Fig. 8C
); apex of laterotergites 8 and 9 acute (
Fig. 8D
) ..........................................................
C. lenti
Grazia, 1968
- Apex of hypandrium with 1+1 narrow, rectangular process (
Fig. 15A
); apex of laterotergites 8 and 9 obtuse (
Fig. 16A
) .................................
C. costaricensis
sp.n
.
8 Humeral angles outlined in black ...................................................................... 9
- Humeral angles not outlined in black.............................................................. 10
9 Dorsal expansions of hypandrium broad, with 1 +1 laminar processes with quadrangular form along dorsal surface, near apex (
Fig. 17A
) ......
C. caxiuanensis
sp.n
.
- Dorsal expansion of hypandrium narrow. Without process on dorsal surface (
Fig. 4E
)...............................................................
C. rolstoni
Grazia-Vieira, 1973
10 Ventral process of hypandrium digitiform (
Fig. 8F
)........
C. dollingi
Grazia, 1987
- Ventral process of hypandrium bilobate (
Fig. 8G
) ........................................... 11
11 Ventral surface of hypandrium mesially bearing 1+ 1 laminar process, oblique to the longitudinal axes of the hypandrium (
Fig. 8G
); parameres short, not reaching the margin of pygophore (
Fig. 9A
); posterior margin of gonocoxites 8 strongly angulate (
Fig. 9B
)..................................
C. tucuruiensis
Grazia & Teradaira, 1980
- Ventral surface of hypandrium without mesially process (
Fig. 9C
); parameres long, almost surpassing the margin of pygophore (
Fig. 9D
); posterior margin of gonocoxites 8 slightly convex..............
C. stysi
Grazia, Schwertner & Greve, 2008
12 Width at base of gonocoxites 9 almost 2.5 times the width at apex; laterotergites 8 with triangular apices (
Fig. 14A
); males unknown ............................................ ..........................................................................
C. luteipennis
(
Westwood, 1837
)
- Width at base of gonocoxites 9 nearly two times the width at apex; laterotergites 8 strongly acute at apices, almost spine like (
Fig. 9E
); pygophore strongly excavated dorsally, reducing the dorsal wall of pygophore to nearly half of the total length of pygophore; parameres long, surpassing segment Xin length (
Fig. 9F
) ................. ............................................................................................
C. vigens
(
Stål, 1860
)