<document id="75FD89D48D1A902A5C2925FE420177AD" ID-DOI="10.12651/JSR.2013.2.2.099" ID-ISSN="2713-8615" ID-Zenodo-Dep="13144615" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722438108010" checkinUser="felipe" docAuthor="Blakemore, Robert J." docDate="2013" docId="03CA8049FFABFFF13626FC3A5C3DFE46" docLanguage="en" docName="JSpeciesRes.2.2.99-126.pdf" docOrigin="Journal of Species Research 2 (2)" docSource="http://dx.doi.org/10.12651/jsr.2013.2.2.099" docStyle="DocumentStyle:4D398BFF45FF8CA9E08F546BB06AE372.2:JSpeciesRes.2012-.journal_article" docStyleId="4D398BFF45FF8CA9E08F546BB06AE372" docStyleName="JSpeciesRes.2012-.journal_article" docStyleVersion="2" docTitle="Amynthas carnosus subsp. carnosus" docType="treatment" docVersion="2" lastPageNumber="101" masterDocId="FFF3F831FFA9FFF33510FD3B5831FFFC" masterDocTitle="Megascolex (Perichaeta) diffringens Baird, 1869 and Pheretima pingi Stephenson, 1925 types compared to the Amynthas corticis (Kinberg, 1867) and A. carnosus (Goto &amp; Hatai, 1899) species-groups (Oligochaeta: Megadrilacea: Megascolecidae)" masterLastPageNumber="126" masterPageNumber="99" pageNumber="101" updateTime="1722438559216" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="E313DAD0F56DF93F1BE699C3AC4D7ED5">Megascolex (Perichaeta) diffringens Baird, 1869 and Pheretima pingi Stephenson, 1925 types compared to the Amynthas corticis (Kinberg, 1867) and A. carnosus (Goto &amp; Hatai, 1899) species-groups (Oligochaeta: Megadrilacea: Megascolecidae)</mods:title>
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<mods:namePart id="2849651A3987240C978C136279EBF333">Blakemore, Robert J.</mods:namePart>
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<taxonomicName id="4C634ADCFFABFFF13626FC3A5D5CFEE5" authority="(Goto &amp; Hatai, 1899)" baseAuthorityName="Goto &amp; Hatai" baseAuthorityYear="1899" box="[822,1389,257,281]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="2" pageNumber="101" phylum="Annelida" rank="subSpecies" species="carnosus" subSpecies="carnosus">
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<emphasis id="B917ED4DFFABFFF13626FC3A5C48FEE4" bold="true" box="[822,1145,257,281]" italics="true" pageId="2" pageNumber="101">Amynthas carnosus carnosus</emphasis>
(
<bibRefCitation id="EFF24CAEFFABFFF13195FC3A5D57FEE5" author="Goto, S. &amp; S. Hatai" box="[1157,1382,257,281]" pageId="2" pageNumber="101" pagination="13 - 24" refId="ref15374" refString="Goto, S. and S. Hatai. 1899. New or imperfectly known species of earthworms. No. 2. Annotations Zoologicae Japonensis 3 (1): 13 - 24." type="journal article" year="1899">Goto &amp; Hatai, 1899</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C37962D4FFABFFF13626FC1A5C3DFE46" pageId="2" pageNumber="101" type="description">
<paragraph id="8BDC315FFFABFFF13626FC1A5B96FEC5" blockId="2.[822,1427,257,442]" box="[822,935,289,313]" pageId="2" pageNumber="101">
(
<figureCitation id="13582DDAFFABFFF1362EFC1A5BAFFEC5" box="[830,926,289,313]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="3.[159,192,1704,1725]" captionStartId-1="4.[157,190,1302,1323]" captionTargetBox-0="[265,1286,263,1647]" captionTargetBox-1="[160,766,262,1270]" captionTargetId-0="figure-104@3.[232,1367,260,1726]" captionTargetId-1="figure-579@4.[151,811,238,1292]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Fig. 2. A. carnosus - A. Korean specimen from Geoman (NIBR IV261234 - DNA w37). B. Japanese neotype of A. carnosus (Tokyo An- 435 from Blakemore, 2012a). These two specimens are almost identical morphologically allowing genetic comparison(Fig. 1)." captionText-1="Fig. 3. A. carnosus - sketch of Hikone specimen (Tokyo An-460 - DNA JET-112) showing ventral view of anterior and dorsal views of prostomium, clitellum and flared male pores posteriorly (other specimen An-461 and JET samples 113-115 agreed 100%). This specimen resembles P. pingi (cf Fig. 11) but its DNA agrees with A. carnosus (as in Fig. 1)." figureDoi-0="http://doi.org/10.5281/zenodo.13144619" figureDoi-1="http://doi.org/10.5281/zenodo.13144623" httpUri-0="https://zenodo.org/record/13144619/files/figure.png" httpUri-1="https://zenodo.org/record/13144623/files/figure.png" pageId="2" pageNumber="101">Figs. 2-3</figureCitation>
)
</paragraph>
<paragraph id="8BDC315FFFABFFF13626FC7A5C3DFE46" blockId="2.[822,1427,257,442]" pageId="2" pageNumber="101">
Delailed by 
<bibRefCitation id="EFF24CAEFFABFFF136A5FC7A5C40FEA5" author="Kobayashi, S." box="[949,1137,321,345]" pageId="2" pageNumber="101" pagination="115 - 138" refId="ref15652" refString="Kobayashi, S. 1936. Distribution and some external characteristics of Pheretima (Ph.) carnosa (Goto et Hatai) from Korea. Science Report of the Tohoku Imperial University 11 (1): 115 - 138." type="journal article" year="1936">Kobayashi (1936)</bibRefCitation>
, 
<bibRefCitation id="EFF24CAEFFABFFF1316BFC7A5D2AFEA5" author="Ohfuchi, S." box="[1147,1307,321,345]" pageId="2" pageNumber="101" pagination="31 - 136" refId="ref15840" refString="Ohfuchi, S. 1937. On the species possessing four pairs of spermathecae in the genus Pheretima, together with the variability of some external and internal characteristics. Saito Ho-On Kai Museum Research Bulletin 12: 31 - 136." type="journal article" year="1937">Ohfuchi (1937)</bibRefCitation>
and 
<bibRefCitation id="EFF24CAEFFABFFF1305DFC7A5BEFFE86" author="Blakemore, R. J." pageId="2" pageNumber="101" pagination="35 - 43" refId="ref14125" refString="Blakemore, R. J. 2012 a. Amynthas carnosus (Goto &amp; Hatai, 1899) redescribed on its neotype (Oligochaeta: Megadrilacea: Megascolecidae). Journal of Species Research 1 (1): 35 - 43 [Available from: www. nibr. go. kr / download? attach _ path = journal &amp; attach = 2012 vol 1 no 1 _ 4. pdf]." type="journal article" year="2012">Blakemore (2012a</bibRefCitation>
; 
<bibRefCitation id="EFF24CAEFFABFFF136FCFC595C05FE86" author="Blakemore, R. J." box="[1004,1076,354,378]" pageId="2" pageNumber="101" pagination="55 - 69" refId="ref14353" refString="Blakemore, R. J. 2013 a. Ulleung-do earthworms - Dagelet Island revisited. Journal of Species Research 2 (1): 55 - 69 [Available from: www. nibr. go. kr / eng / event / journal _ spe _ 3. jsp]." type="journal article" year="2013">2013a</bibRefCitation>
; 
<bibRefCitation id="EFF24CAEFFABFFF13152FC5A5CB8FE85" author="Blakemore, R. J." box="[1090,1161,353,377]" pageId="2" pageNumber="101" pagination="15 - 54" refId="ref14406" refString="Blakemore, R. J. 2013 b. Jeju-do earthworms (Oligochaeta: Megadrilacea) - Quelpart Island revisited. Journal of Species Research 2 (1): 15 - 54 [Available from: www. nibr. go. kr / eng / event / journal _ spe _ 3. jsp]." type="journal article" year="2013">2013b</bibRefCitation>
) (cf 
<taxonomicName id="4C634ADCFFABFFF131D2FC595D2CFE85" authorityName="Stephenson" authorityYear="1925" box="[1218,1309,354,377]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="2" pageNumber="101" phylum="Annelida" rank="species" species="pingi">
<emphasis id="B917ED4DFFABFFF131D2FC595D2CFE85" box="[1218,1309,354,377]" italics="true" pageId="2" pageNumber="101">A. pingi</emphasis>
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described below). DNA data (
<figureCitation id="13582DDAFFABFFF1310EFCB95C6AFE66" box="[1054,1115,386,410]" captionStart="Fig" captionStartId="2.[157,190,1924,1945]" captionTargetBox="[199,1380,1077,1894]" captionTargetId="figure-304@2.[190,1400,1065,1897]" captionTargetPageId="2" captionText="Fig. 1. Phylotree from MEGA5 program Maximum Likelihood defaults - sample codes explained in the Appendix; ◆- samples have figured specimens; ◇- ex GenBank. Appears to show significant genetic difference between specimens superficially compliant with broad A. corticis definition as discussed below that taxon herein. Synonym priority for either Gp 1 or Gp 2 is currently unresolved." figureDoi="http://doi.org/10.5281/zenodo.13144617" httpUri="https://zenodo.org/record/13144617/files/figure.png" pageId="2" pageNumber="101">Fig. 1</figureCitation>
) indicates a new sub-species as next described.
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