A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data Author Tarasov, Sergei text Zootaxa 2017 2017-10-04 4329 2 101 149 journal article 31902 10.11646/zootaxa.4329.2.1 7e33ec7d-211b-4042-8d07-344364ee805d 1175-5326 1001915 8226E27D-E3A9-481A-B51B-558643Bbb03A Parachorius semsanganus Tarasov & Keith, 2011 ( Figs. 22 , 27D ) Parachorius semsanganus Tarasov & Keith, 2011 : 51 Type locality . Phou Sane Mountain, northern Laos. Distribution. Northern Laos , Phou Sane Mountain. Diagnosis. This species stands out in being the largest representative of Parachorius . It can be separated from all other species of Parachorius by metatibia with large teeth in males and inner margin denticulate and parameres with apex bent upward and membranous lobes. Description. Body. Body black, length 8.2–10.6 mm . Elytra uniformly colored. Head. Clypeus with two teeth, outer margin of each tooth very slightly notched basally. Pronotum. Pronotum rounded laterally. Frontal angles rounded. Disc covered with punctures separated by 1–2 puncture diameter(s). Pronotum in anterior-posterior direction notably shorter than elytra. Elytra. Elytra covered with sparse punctures. Wings. Wings normally developed. Legs. Profemur with almost straight or sinuate anterior margin. Protibia with apical spur reaching protarsomere 2. First tooth notably modified in males, approximately twice as wide as second tooth; first and second teeth distinctly separated from each other; first tooth distinctly separated from denticles located between first and second teeth. Metafemoral posterior margin serrate (sometimes slightly), simple, not modified. Metatibia slightly sinuate, inner margin denticulate with large teeth on inner margin, produced with a small angle apically. Metasternum. Transverse basal ridge of metasternum reduced and extremely short. Aedeagus. Parameres symmetrical, bearing membranous lobes and bending upward apically. Material examined. Holotype : Laos : 1 male , " LAOS-NE , Xieng Khouang prov., 19°38.20'N 103°20.20'E , Phonsavan ( 30 km NE) : PHOU SANE Mt. , 1420 m , 10.-30.v.2009 , D. Hauck leg. / HMB Basel , NMPC Prague Laos 2009 Expedition : M. Brancucci, M. Geiser, Z. Kraus , D. Hauck , V. Kubán / HOLOTYPE Parachorius semsanganus D. Keith & S. Tarasov det. 2011", 19°38.2'N , 103°20.2'E ( NHMB ) . Paratypes : 4 males , 3 females , Xieng Khouang Province, 30 km NE Phonsavan : Ban Na , Lam Phou Sane Mountain , 19°38'N , 103°20'E , 1300– 1500 m , on 10–30/v/2009 ( M. Brancucci ) ( NHMB ) . Paratypes : 2 males , Xieng Khouang Province, Phonsavan ( 30 km NE): Phou Sane Mountain , Laos 2009 Expedition : M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubán, 19°38.2'N , 103°20.2'E , 1420 m , on 10–30/v/2009 ( Z. Kraus ) ( NHMB ) . Paratypes : 9 females , Xieng Khouang Province, Phonsavan ( 30 km NE): Phou Sane Mountain , Laos 2009 Expedition : M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubán, 19°38.2'N , 103°20.2'E , 1420 m , on 10–30/v/2009 ( D. Hauck ) ( NHMB ) .