Systematics of the green lacewing tribe Ankylopterygini Navás, 1910 (Neuroptera: Chrysopidae: Chrysopinae) from China Author Wu, Jingyu 0000-0001-6677-7726 Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659 wujingyu176@163.com Author Liu, Xingyue 0000-0001-6677-7726 Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659 wujingyu176@163.com text Zootaxa 2024 2024-11-22 5540 1 1 169 http://dx.doi.org/10.11646/zootaxa.5540.1.1 journal article 10.11646/zootaxa.5540.1.1 1175-5334 14240496 4DBD3A92-F14B-4C5B-95B3-2B430EC197DD [57] Retipenna variegata Brooks, 1986 ( Figs 67 , 87 ) Retipenna variegata Brooks, 1986: 67 . Type locality: India (Khasi Hills). Holotype in NHML. Diagnosis. This species is characterized by the head with spots on the gena, the thorax without markings, the dark brown veins in basal areas and gradate series of fore- and hindwings, the median arch of male gonarcus with shallow U-shaped invagination, the long narrow arcessus rounded at base and pointed at tip. Description. Male. Body length 11.8 m , forewing length 14.7 mm , hindwing length 13.3 mm . Head yellow, vertex raised. Compound eyes prominent and black. Antennae yellow, scape and pedicel wider than flagellum. Frons and clypeus not marked. Gena marked black spots. Maxillary and labial palpi brownish yellow. Pronotum anterior margin shorter than posterior margin, mesonotum and metanotum without markings. Legs with dense setae. Pretarsal claws brown, with basal dilation. Forewing slightly narrow, dark brown veins present in three areas: veins in basal area, inner gradate series and apical radial crossveins, outer gradates and forking marginal veins. Pterostigma pale brown. Costal area with 20–25 crossveins; 12–15 radial crossveins present. m1 smaller than m2 ; im ovoid; 10–11 psm-psc crossveins present. c1 smaller than c2 , dcc closed. A1 forked, A2 and A3 simple. Inner gradate series with 6–7 crossveins, outer gradate series with 8 crossveins. Hindwing narrow, apical radial crossveins and gradate series brown. Costal area with 15–17 crossveins; 13–14 radial crossveins and 8 psm-psc crossveins present. Inner gradate series with 6–7 crossveins, outer gradate series with 7 crossveins. Abdomen with dense setae. Tergum VII and sternum VII subquadrate in lateral view; tergum VIII slightly shorter than tergum VII; sternum VIII fused with sternum IX and with small tubercle near tip; tergum IX fused with ectoproct. Callus cerci ovoid. Gonarcus arcuate, median arch with shallow U-shaped invagination, lateral arms narrow at base and wide at tip; entoprocessus widest at tip; arcessus long, rounded at base and pointed at tip. Female. Body length 10.0 mm, forewing length 16.0 mm, hindwing length 14.0 mm. Sternum VII longer than tergum VII; tergum VII subquadrate in lateral view; tergum VIII shorter than tergum VII; tergum IX fused with ectoproct. Callus cerci ovoid. Subgenitale bilobed apically; spermatheca thick, vela extremely short, ventral impression deep, duct long and coiled. FIGURE 67. Retipenna variegata Brooks, 1986 . A. Dorsal habitus; B. Head, frontal view; C. Head, lateral view; D. Thorax, dorsal view; E. Male genitalia, lateral view; F. Gonarcal complex, dorsal view; G. Gonarcal complex, lateral view; H. Arcessus; I. Female genitalia, lateral view; J. Subgenitale; K. Spermatheca. Scale bar: 5 mm (A), 1 mm (B–D). Material. 1♂ , China , Yunnan , Yingjiang , Hulukou , 2018.IV ( CAU ) ; 1♀ , China , Xizang , Mêdog , 777.9 m , 2019.V.30 , Rongrong Shen & Yingnan He ( CAU ). Distribution. China ( Xizang , Yunnan ); India ( Khasi Hills ). Remarks. Brooks (1986) established Retipenna variegata and provided genital information. This species is firstly recorded from China and redescribed.