The apterous endemic genus Omphra Dejean (Coleoptera: Carabidae: Helluonini) of the Indian subcontinent: taxonomy with notes on habits and distributional patterns Author Raj, Shiju T. Post Graduate and Research Department of Zoology The New College, Royapettah Chennai- 670014, Tamil Nadu, INDIA shijutraj@gmail.com Author Sabu, Thomas K. Litter Entomology Research Unit Post Graduate and Research Department of Zoology St. Joseph’s College, Devagiri Calicut- 673008, Kerala, INDIA sabukthomas@gmail.com Author Danyang, Zhao Zhao Danyang Department of Entomology College of Natural Resources and Environment South China Agricultural University, Wushan Guangzhou- 510640, CHINA text Insecta Mundi 2012 2012-02-29 2012 206 1 15 journal article 7452 10.5281/zenodo.4670934 0c820dc3-bd71-4eba-902e-59bd37283060 1942-1354 4670934 Key to the Indian genera of the tribe Helluonini (modified from Sloane 1914 ) 1. Wingless forms; mentum with a strongly developed stout median tooth; legs stout. ................. ......................................................................................................................... Omphra Dejean – Winged forms; mentum with an elongated spiniform median tooth; legs slender. ................... 2 2(1). Tarsi with fourth joint deeply bilobed. ............................................................ Creagris Nietner – Tarsi with fourth joint entire. ...................................................................................................... 3 3(2). Fourth maxillary palpomere securiform. ....................................................... Colfax Andrewes – Fourth maxillary palpomere not securiform. ............................................ Macrocheilus Hope