Order Rodentia - Family Echimyidae Author Wilson, Don E. Author Reeder, DeeAnn text 2005 The Johns Hopkins University Press Baltimore Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2 1575 1592 book chapter 0-8018-8221-4 10.5281/zenodo.7316535 Phyllomys nigrispinus Wagner 1842 Phyllomys nigrispinus Wagner 1842 , Arch. Naturgesch., 1: 361 . Type Locality: Brazil , São Paulo , Floresta Nacional de Ipanema, 20 km NW Sorocaba ( 23 ^o 26’S , 47 ^o 37’W ); as amended by Emmons et al. (2002) . Vernacular Names: Blacked-spined Atlantic Tree-rat . Distribution: SE Brazil , from the state of Rio de Janeiro to Paraná , mainly along the coastal zone, but extending inland to W São Paulo to at least 850 m . Conservation: IUCN – Lower Risk (lc) as Echimys nigrispinus . Discussion: Karyotype has 2n=52 ( Leite, 2003 ).