The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae) Author Ramahandrison, Andriamirado Tahina https://orcid.org/0000-0002-0833-8730 Departement de Biologie et Ecologie Vegetales, Faculte des Sciences, BP 906, Universite d'Antananarivo, Antananarivo, Madagascar & Sorbonne Universite, Institut de Systematique, Evolution, Biodiversite (UMR 7205), MNHN SU CNRS EPHE UA, Case 05, 7 quai St Bernard, Paris, France Author Rakouth, Bakolimalala https://orcid.org/0000-0001-5710-2006 Departement de Biologie et Ecologie Vegetales, Faculte des Sciences, BP 906, Universite d'Antananarivo, Antananarivo, Madagascar Author Manuel, Michael Sorbonne Universite, Institut de Systematique, Evolution, Biodiversite (UMR 7205), MNHN SU CNRS EPHE UA, Case 05, 7 quai St Bernard, Paris, France manuel1570@yahoo.fr text ZooKeys 2022 2022-11-02 1127 1 60 http://dx.doi.org/10.3897/zookeys.1127.85737 journal article http://dx.doi.org/10.3897/zookeys.1127.85737 1313-2970-1127-1 4759AFC32EFD47A7853F645FB32829BA D72971CE12A85992AFFD69F186474E85 Laccophilus luctuosus Sharp, 1882 Type locality. Madagascar. Material examined. 1 ♂ , 1 ♀ : MAK-21. Distribution. Madagascar (widespread in lowlands) ( Guignot 1959-1961 ; Bertrand and Legros 1971 ; Rocchi 1991 ; Bistroem et al. 2015 ). Habitat in study area. This species was encountered only once in a peripheral site located in an open area. The habitat was a small isolated temporary puddle with sandy bottom and clear water under Phragmites , on the west bank of the Makaikely River. There was a moderate amount of plant debris, and no vegetation.