On the identity of species of the huntsman spider genus Thelcticopis Karsch, 1884 (Araneae: Sparassidae: Sparianthinae) from India, Myanmar, Pakistan, and Sri Lanka
Author
Sankaran, Pradeep M.
Division of Arachnology, Department of Zoology, Sacred Heart College, Thevara, Cochin, Kerala 682 013, India.
Author
Sherwood, Danniella
Arachnology Research Association, 124 City Road, London, ECIV 2 NX, United Kingdom. & Fundación Ariguanabo, 4111, Calle 58, e / ave. 41 y ave. 43, San Antonio de los Baños, Provincia Artemisa c. p. 18100, Cuba.
Author
Jäger, Peter
Arachnology, Senckenberg Research Institut, Mertonstrasse 17 - 21, 60325 Frankfurt am Main, Germany.
text
Zootaxa
2024
2024-06-06
5463
3
301
338
http://dx.doi.org/10.11646/zootaxa.5463.3.1
journal article
298429
10.11646/zootaxa.5463.3.1
5390ab23-704a-4644-8307-4bfc73751bcf
1175-5326
11611277
6E261F38-7196-4C87-AE81-E09996D055F4
Thelcticopis birmanica
Thorell, 1895
stat. rev.
Figs 6–7
,
26
Thelcticopis birmanica
Thorell, 1895: 274
(
♀
).
Pocock 1900: 271
(
♀
).
Thelcticopis canescens
Simon
:
Gravely 1931: 246
, ad part.: fig. 7B [doubtful identity] (
♀
).
Type material.
Holotype
♀
,
MYANMAR
:
Tanintharyi
:
Tenasserim
[ca.
12°05’7.3’’N
,
99°0’40.95’’E
;
47 m
a.s.l.
], date unknown,
E.W. Oates
leg., old type number: 1971/212 (
NHMUK
1895.9.21.867; examined).
Diagnosis.
Females of
T
.
birmanica
stat. rev.
resemble
T. canescens
Simon, 1887
,
T. folia
Jäger & Praxaysombath, 2009
: fig. 106,
T. karnyi
Reimoser, 1929
: fig. 3,
T. kirankhalapi
Ahmed
et al.,
2015
: figs 4A–B,
T. klossi
Reimoser, 1929
: fig.2,
T. virescens
Pocock, 1901
(
Figs 22D–E
,
24C
herein) and
T.zhengi
Liu, Li & Jäger, 2010
(
Liu
et al.
2010
: fig. 4F) by the continuous anterior epigynal margin (
Figs 6C
,
7A
). The species can be distinguished from
T. folia
,
T. karnyi
,
T. kirankhalapi
,
T. klossi
and
T. zhengi
by its median septum 1.5 times longer than wide (vs. as long as wide or slightly wider than long in
T. folia
and
T. zhengi
, and vs. 2.2 to 4 times longer than wide in
T. karnyi
,
T. klossi
and
T. virescens
). In
T. kirankhalapi
, the median septum is 1.75 times longer than wide, but it can be easily distinguished by its vulva having a simple duct system running from anterior to posterior broad over its entire length, and by its median septum roughly of the same width over its entire length, whereas
T. birmanica
has narrow ducts in the anterior half, a bend posteriorly, and a median septum narrowing abruptly in the posterior half (cf.
Figs 6C–D
,
7B
vs.
Ahmed
et al.
2015
: figs 4A–B). The species can be distinguished from
T. canescens
by having an abruptly narrowing median septum in posterior half (continuously tapering in
T. canescens
).
FIGURES 4A–G.
Thelcticopis bicornuta
Pocock, 1901
, holotype male. A–B Prosoma (A dorsal, B ventral); C Opisthosoma, dorsal; D–F Left palp (D prolateral, E ventral, F retrolateral); G Labels in the holotype vial. Scale bars: A–F, 1 mm. Photographs by Danniella Sherwood, © The Trustees of the Natural History Museum, London.
FIGURES 5A–D.
Thelcticopis bicornuta
Pocock, 1901
, holotype male. A–B Left palp (A ventral, B retrolateral); C Eye arrangement, dorsal; D Right chelicera, ventral. Abbreviations: C—conductor; dRTA—dorsal part of retrolateral tibial apophysis; dSe—distal setae; E—embolus; Se—bunch of setae; Sp—spermophor; VDL—ventro-distal lobe; vRTA—ventral part of retrolateral tibial apophysis; TA—tegular apophysis. Drawings by Peter Jäger.
Supplementary description.
Female
(
holotype
;
Figs 6A–B
,
7D–E
). Colouration: brown. [after
Thorell 1895
: carapace red-brown, clothed with grey setae (partly rubbed off); opisthosoma yellow-brown]. Thoracic striae indistinct. Fovea deep, longitudinal, straight. Chelicerae with three promarginal and seven retromarginal teeth (
Fig. 7E
). Body length 17.5. Carapace 6.5 long. Opisthosoma 8.0 long, 6.38 wide. Chelicerae 1.76 long. Eye sizes and interdistances (
Fig. 7D
): AME 0.40, ALE 0.33, PME 0.27, PLE 0.26; AME–AME 0.43, AME–ALE 0.57, AME– PME 0.45, ALE–PLE 0.58, PME–PME 0.90, PME–PLE 0.88. Legs: I 16.70 (5.12, 2.21, 4.30, 3.86, 1.21), II 16.31 (5.09, 2.25, 4.10, 3.63, 1.24), III 13.67 (4.41, 2.31, 2.77, 2.78, 1.40), IV 18.54 (5.84, 2.27, 4.35, 4.55, 1.53). Leg formula: 4123. Spination: legs: femur I–III 323, tibia I–II 000(10), III–IV 0006, metatarsus I–II 0002, III–IV 0006.
Genitalia
(
Figs 6C–D
,
7A–C
): epigynal field longer than wide, with tiny, anterior bands attached to the anterior margin and one slit sensillum close to the field on the right side (
Figs 6C
,
7A
). Lateral lobes short, i.e., roughly one third of epigynal field length, sclerotized, widely separated, with posteriad posterior projections (
Figs 6C
,
7A
). Median septum with truncated posterior part (
Figs 6C
,
7A
). Copulatory ducts sclerotized, widely separated and with spherical structures in anterior half (
Figs 6D
,
7B–C
). Spermathecae large and almost touching each other in posterior half (
Figs 6D
,
7B
). Fertilisation ducts short and narrow, tips laterad to slightly antero-laterad (
Figs 7B–C
).
FIGURES 6A–E.
Thelcticopis birmanica
Thorell, 1895
stat. rev.
, holotype female. A–B Habitus (A dorsal, B ventral); C Epigyne, ventral; D Vulva, dorsal; E Labels in the holotype vial. Scale bars: A–D, 1 mm. Photographs by Danniella Sherwood, © The Trustees of the Natural History Museum, London.
FIGURES 7A–E.
Thelcticopis birmanica
Thorell, 1895
stat. rev.
, holotype female. A Epigyne, ventral; B–C Vulva (B dorsal, C anterior); D Eye arrangement, dorsal; E Left chelicera, ventral. Abbreviations: AB—anterior band of epigynal field; CD— copulatory duct; EF—epigynal field; FD—fertilisation duct; LL—lateral lobe; MS—median septum; S—spermatheca; SS—slit sensillum. Drawings by Peter Jäger.
Male
. Unknown.
Distribution.
Myanmar
(
Tanintharyi
) (
Thorell 1895
;
Gravely 1931
) (
Fig. 26
).
Remarks.
Thorell (1895)
described this species based on a female specimen collected from Tenasserim (=
Tanintharyi
),
Burma
(=
Myanmar
).
Gravely (1931)
, with certain hesitation, suggested a possible synonymy of this species with
T. canescens
, though he did not formally establish it, but this synonymy was subsequently accepted by
Roewer (1955)
. Examination of the
holotype
of
T. birmanica
revealed that its epigyne is different from that of
T. canescens
(as shown in
Gravely 1931
: fig. 7C based on a female from Tavoy, the type locality of
T. canescens
), a species described from
Myanmar
by
Simon (1887b)
; as already noted by
Pocock (1900)
and
Gravely (1931)
the epigyne of
T. birmanica
presents an epigynal plate with a broad, median cleft accommodating a middle piece with truncated apex
vs.
narrow, median cleft accommodating a middle piece with rounded apex (cf.
Figs 6C
,
7A
vs.
Gravely 1931
: fig. 7B). Moreover,
Pocock (1900)
mentioned that
T
.
birmanica
is smaller in size than
T
.
canescens
. As a consequence, we remove
T. birmanica
stat. rev.
from the synonymy of
T. canescens
and consider it as a valid species.