A monograph of the Xyleborini (Coleoptera, Curculionidae, Scolytinae) of the Indochinese Peninsula (except Malaysia) and China
Author
Smith, Sarah M.
Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA
https://orcid.org/0000-0002-5173-3736
camptocerus@gmail.com
Author
Beaver, Roger A.
161 / 2 Mu 5, Soi Wat Pranon, T. Donkaew, A. Maerim, Chiangmai 50180, Thailand
Author
Cognato, Anthony I.
Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA
text
ZooKeys
2020
983
1
442
http://dx.doi.org/10.3897/zookeys.983.52630
journal article
http://dx.doi.org/10.3897/zookeys.983.52630
1313-2970-983-1
7DED4CE2934C4539945F758930C927F9
C890C7FD4B2D57A8B1A062305ED42D53
Amasa Lea, 1894
Amasa
Lea, 1894: 322.
Pseudoxyleborus
Eggers, 1930: 206. Synonymy:
Wood 1984
: 223.
Anaxyleborus
Wood, 1980: 90. Synonymy:
Wood 1983
: 647.
Type species.
Amasa thoracica
Lea, 1894 =
Tomicus truncatus
Erichson, 1842; monotypy.
Diagnosis.
2.5-5.0 mm, 2.11-3.4
x
as long as wide.
Amasa
is distinguished by the declivity truncate, margined with a circumdeclivital ring; antennal club flattened, types 4 or 5 (typically type 4), club sutures sinuate, two sutures visible on posterior face; protibiae typically slender, inflated and granulate on posterior face (rarely distinctly triangular or unarmed on posterior face); anterior margin of pronotum with a row of serrations; scutellum flat, flush with elytral surface; declivital face with three striae; procoxae contiguous or narrowly separated; and mycangial tufts absent.
Similar genera.
Cyclorhipidion
,
Pseudowebbia
,
Truncaudum
,
Webbia
,
Xylosandrus
.
Distribution.
Distributed throughout Asia and Australasia, also occurring in Madagascar. One species has been introduced to Brazil, Chile and Uruguay (
Flechtmann and Cognato 2011
;
Gomez
et al. 2017
;
Kirkendall 2018
).
Gallery system.
This usually comprises a short radial tunnel leading to a single, large, flat brood chamber, extending in the longitudinal plane.
Remarks.
Amasa
is easily confused with other species possessing truncate declivities in the genera listed above. Most species can be readily distinguished by the type 4 antennal club with sinuate sutures and the presence of only three striae on the declivital face.
Previous morphological studies of
Amasa
have suggested that species are very morphologically variable (
Hulcr and Cognato 2013
). As a result, many species were considered conspecific and part of a morphological continuum. Molecular data generated as part of this study has demonstrated that
Amasa
species are actually morphologically conserved even across broad ranges (
Smith et al. 2020
).
Amasa
species outside our coverage area are thus in need of revision. Potentially much of the diversity is awaiting discovery.