World reclassification of the Cardiophorinae (Coleoptera, Elateridae), based on phylogenetic analyses of morphological characters Author Douglas, Hume B. text ZooKeys 2017 655 1 130 http://dx.doi.org/10.3897/zookeys.655.11894 journal article http://dx.doi.org/10.3897/zookeys.655.11894 1313-2970-655-1 8C475FAB25E044CEA2FBC3B83F316D8C 8C475FAB25E044CEA2FBC3B83F316D8C Ryukyucardiophorus Ohira , 1973 Figs 161-162 Ryukyucardiophorus Ohira , 1973a: 32. Type species: Paracardiophorus loochooensis , Miwa 1934 : 255. Prothorax. Pronotum with carina extending anterad from hind angles following lateral edge (ie, not below lateral edge of prothorax, but in some species not distinguishable from the dorsal hind angle carina), not reaching anterior edge. Legs. Protibiae not modified for digging; tarsi with tarsomere 4 not lobed or lamellate, claws with 2 points per side. Bursa copulatrix with paired proximal (largest) sclerites bilobed with attached semi-sclerotised membrane with spines (Fig. 162). Also. Procoxal cavities apparently closed; aedeagus parameres apices without lateral expansions. Known from Japan, and Taiwan, 4 spp.