A new species of Helgreelia Gaimari (Diptera: Odiniidae) from Brazil, with a key to the Neotropical species of Odiniidae Author Filho, Fernando Da Silva Carvalho Author Esposito, Maria Cristina Author Santos, Rita De Cássia Oliveira Dos text Zootaxa 2009 2219 61 68 journal article 10.5281/zenodo.189998 a74fab7c-4b46-4758-9b71-66af304dd1ad 1175-5326 189998 Key to the Neotropical species of Odiniidae (modified from Shewell (1960) , Sabrosky (1969), Prado (1973) and Gaimari (2007)) 1. Posterior ocelli in front of line between medial vertical setae; ocelli usually arranged in isosceles triangle with posterior ocelli closely spaced; frons usually relatively arched and sometimes conspicuously tuberculate; scutellar disc sometimes setulose; in profile, face and frons more or less in line or sloping; TRAGINOPINAE ............................ 7 - Posterior ocelli in line with medial vertical setae; ocelli arranged in equilateral triangle or with distance between posterior pair greater than between either posterior ocelli more widely spaced; frons flat to slightly concave and never tuberculate; scutellar disc bare; in profile, face and frons meeting at a angle; ODINIINAE , Odinia Robineau- Desvoidy ....................................................................................................................................................................... 2 2. Wing with few black spots, usually restricted to cross-veins ....................................................................................... 3 - Wing with numerous black spots, most of them surrounded by a whitish corona ..................................................... 4 3. Cell R1 with two transverse black spots................... Odinia biguttata Sabrosky ( Virgin Islands , Puerto Rico , USA ) - Cell R1 without spot ............................................................................................... Odinia brevitibia Shewell ( Brazil ) 4. Oral vibrissa and one or two subvibrissal setae distinctly flattened .... Odinia williamsi Johnson (Galápagos Islands) - Oral vibrissa and subvibrissal setae slender, not unusually broadly flattened.............................................................. 5 5. Antennal segments almost entirely yellow ................ Odinia parvipunctata Sabrosky ( Virgin Islands , Puerto Rico ) - Antennal segments with black or brown spots ............................................................................................................. 6 6. Anepisternum gray with brown stripe ventrally; cell cua1 with one small and one large brown spots, large one tran- versing cell, beginning in vein CuA1 and ending in wing ventral margin ............... Odinia surumuana Prado ( Brazil ) - Anepisternum gray with brown stripe dorsally; cell cua1 with two very small brown spots, close to vein CuA1 ......... .............................................................................................. Odinia coronata Sabrosky ( USA , El Salvador , Panama ) 7. Ocelli on large tubercle well forward of vertex ........................................................................................................... 8 - Ocelli not on tubercle.................................................................................................................................................. 10 8. Arista long-plumose; vein M1 strongly curved forward at apex ..................................... Paratraginops Hendel ….. 9 - Arista pubescent; vein M1 straight ................ Neotraginops clathratus (Hendel) ( Peru , Paraguay , Brazil , Colombia ) 9. Base of scutellum thickly haired. Anterior four fronto-orbital setae not set in fuscous spots........................................ .......................................................................................................... Paratraginops pilicornis Cress ( Guyana , Brazil ) - Base of scutellum bare. Anterior four fronto-orbital setae set in fuscous spots ............................................................. .................................................................................................................. Paratraginops plaumanni Shewell ( Brazil ) 10. Costal vein extended to vein M1 ................................................................................................................................. 11 - Costal vein extended to vein R4+5 or slightly beyond.................................................................................................. 22 11. Scutellum at most bearing 2–3 setulae along margin ......................................................... Lopesiodinia Prado ..... 12 - Scutellar disc and margins bearing numerous setulae ............................................................................................... 13 12. Arista pubescent; front little narrowed posteriorly .............................................. Lopesiodinia diversa Prado ( Brazil ) - Arista with long rays; front very narrowed posteriorly ................................ Lopesiodinia alvarengai Prado ( Brazil ) 13. Only anterior fronto-orbital seta inclinate ........................................................................ Schildomyia Malloch …. 14 - Several anterior fronto-orbital setae inclinate.......................................... Neoschildomyia fusca Gaimari ( Costa Rica ) 14. Arista long-plumose.................................................................................................................................................... 15 - Arista pubescent.......................................................................................................................................................... 16 15. Clipeo with velvet brown spot ................................................................................... Schildomyia lanei Prado ( Brazil ) - Clipeo without velvet brown spot........................................... Schildomyia vittithorax Malloch ( Costa Rica , Panama ) 16. With one postpronotal seta; acrosthical setae absent .................................................................................................. 17 - With two postpronotal setae; one pair of acrosthical seta........................................................................................... 18 17. Lunule with a large velvet black spot .................................................... Schildomyia trinidadensis Hennig ( Trinidad ) - Lunule without a large velvet black spot .......................................................... Schildomyia peruviana Hennig ( Peru ) 18. Scutellum pale-golden pruinose, except a median brownish spot in anterior edge ....................................................... .............................................................................................................................. Schildomyia flavida Prado ( Brazil ) - Scutellum grayish pruinose, with light-brown spots in many parts of scutellum, forming patterns ......................... 19 19. Fronto-orbital setae set on brown spots; antenna with dark spots ............................................................................. 20 - Fronto-orbital setae not set on brown spots; antenna almost entirely yellow................................................................. .................................................................................................................... Schildomyia brevihirta Malloch ( Panama ) 20. Thorax with spots scattered, not forming patterns. Abdomen light-brown. Male fifth tergite reticulate ...................... ........................................................................................................................... Schildomyia reticulata Prado ( Brazil ) - Abdomen black. Thorax with spots arranged in line. Male fifth tergite normal ...................................................... 21 21. Lunule with a velvet black spot; basoflagellomere yellow, without black spots .. Schildomyia goyana Prado ( Brazil ) - Lunule without a velvet black spot; basoflagellomere with black spot on inner and outer surface ............................... ............................................................................................................. Schildomyia punctifrons Malloch ( Costa Rica ) 22. Postocellar setae present; posterior part of head straight from dorsal view ................................................................... ............................................................................................................... Pradomyia hadromera Gaimari ( Costa Rica ) - Postocellar setae absent; posterior part of head concave from dorsal view ............................................................... 23 23. Several anterior fronto-orbital setae inclinate; scutellar disc setulose................ Shewellia agromyzina Hennig ( Peru ) - With single inclinate fronto-orbital seta; scutellar disc bare .............................................. Helgreelia Gaimari …. 24 24. Abdomen orange, with fine pale setulae......................................................... Helgreelia albeto Gaimari ( Costa Rica ) - Abdomen predominately brown, with black setulae ................................................................................................. 25 25. Abdominal tergites wholly brown, without yellow margin; arista entirely pubescent ................................................... ...................................................................................................................... Helgreelia parkeri Gaimari ( Costa Rica ) - Abdominal tergites mostly brown with posterior margin yellow; arista plumose on the basal half ............................. ............................................................................................................................ Helgreelia gaimarii sp. nov. ( Brazil )