A taxonomic review of the genus Euboeus Boieldieu, 1865 (Coleoptera: Tenebrionidae: Helopini) of the Caucasus, Iran and Turkmenistan Author Nabozhenko, Maxim V. text Zootaxa 2022 2022-07-01 5159 4 451 486 journal article 88477 10.11646/zootaxa.5159.4.1 cffc486a-265f-4d35-a289-a4a54845c77d 1175-5326 6785909 CA6AF6AD-9E4A-4BE3-8B52-6D8E5EA3641F Key to species of the genus Euboeus of the Caucasus, Iran and Turkmenistan on females (without E. prometheus , E. arzanovi and E. zubovi whith unknown females) 1 Pronotum and head dorsally strongly shiny, dark-bluish or dark-violet, elytra black ( Fig. 19B )............ E. chorasanicus - Body dorsally dull, completely black ( Figs 2D , 4E , 7E , 9C , 11D , 12B , 13D , 14C , 15C , 17C )......................... 2 2 Puncturation of head and pronotum coarse and dense, punctures subequal or larger than interpuncture distance; sometimes punctures coarse and sparse at middle of pronotum ( Figs 1A , 2E, F , 4F, G , 5D, E , 6B–D , 7F, G )...................... 3 - Puncturation of head and pronotum fine and sparse, puncture diameter much lesser than interpuncture distance ( Figs 9D, E , 10A, B , 11E, F , 13E, F , 14D, E , 15D, E , 17D, E ), at least in anterior half ( Fig. 12B )................................ 6 3 Pronotum with very coarse and dense puncturation of longitudinally elongated punctures, especially on lateral sides of disc; median smooth line of pronotum elevated near base and separated from another surface by impression with very dense merged punctures ( Fig. 2D, E, F ).................................................................... E. corrugosus - Pronotum with coarse and dense puncturation of simple round, not merged punctures; median line not elevated near base or absent ( Figs 4E , 6B–D , 7E )............................................................................. 4 4 Head densely pubescent by reddish, recumbent, long setae ( Figs 6B–D )................................... E. vicinus - Head bare ( Figs 4E , 7E )............................................................................... 5 5 Metaventrite and abdominal ventrites pubescent with long recumbent hairs ( Fig. 4C, D ).................. E. gorganicus - Metaventrite and abdominal ventrites bare, covered with simple very short setae ( Fig. 7C, D )............... E. zoroaster 6 Lateral margins of pronotum not completely beaded, bead widely interrupted at middle ( Figs 14C , 17C )................ 7 - Lateral margins of pronotum completely beaded ( Figs 9C , 11D , 13D ) or completely not beaded ( Fig. 12B , 15C )......... 8 7 Prothoracic hypomera wrinkled ( Fig. 17F )...................................................... E. quadricollis - Prothoracic hypomera coarsely punctured ( Fig. 14F )............................................. E. krivokhatskyi 8 Lateral margins of pronotum not beaded ( Figs 12B , 15C )..................................................... 9 - Lateral margins of pronotum completely beaded ( Figs. 7C, D, E , 9D, E, F , 10D, E, F ).............................. 10 9 Prothoracic hypomera with coarse large longitudinal wrinkles ( Fig. 12F ). Lateral margins of pronotum strongly rounded at middle and widely emarginated at base ( Fig. 12B )................................................. E. kalashiani - Prothoracic hypomera with dense, large, foveolate punctures ( Fig. 15F ). Lateral margins of pronotum weakly evenly rounded ( Fig. 15C )................................................................................... E. merkli 10 Pronotum widest ahead of middle ( Fig. 9C ). Prothoracic hypomera not flattened along margin, with small confused microwrinkles with poorly distinguished sparse smoothed puncturation against their background; wrinkles with microgranulation at anterior third ( Fig. 9F ).......................................................................... E. grimmi - Pronotum widest at middle or behind middle ( Figs 11D , 13D ). Prothoracic hypomera narrowly ( Fig. 13G ) or widely ( Fig. 11G ) flattened along margin, coarsely and densely punctured ( Fig. 13G ) or coarsely longitudinally wrinkled ( Fig. 11G ), without microgranulation.................................................................................... 11 11 Prothoracic hypomera with coarse longitudinal wrinkles; lateral margin of prothoracic hypomera widely flattened ( Fig. 11G )......................................................................................... E. huedepohli - Prothoracic hypomera with coarse puncturation and smooth wrinkles at middle; lateral margin of prothoracic hypomera very narrowly weakly flattened ( Fig. 13G )............................................................ E. kasatkini