Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species
Author
Budha, Prem B.
Author
Naggs, Fred
Author
Backeljau, Thierry
text
ZooKeys
2017
675
129
156
http://dx.doi.org/10.3897/zookeys.675.13252
journal article
http://dx.doi.org/10.3897/zookeys.675.13252
1313-2970-675-129
E5C8F163D61547B98418CEE8D71A7DAB
Rishetia cf. mastersi Godwin-Austen, 1920
Figs 2F, 8A
Glessula
(Rishetia) mastersi
Godwin-Austen, 1920:
Godwin-Austen 1920
, p. 46, pl. 161, fig. 14., pl. 162, fig. 22.
Glessula mastersi
Godwin-Austen, 1920:
Ramakrishna et al. 2010
, p. 166.
Material examined.
CDZMTU065/5 shells, Kurintar, Chitwan, degraded riverine bushes with big boulders, mixed
Shorea robusta
forest, 420-527 m, 27.875820N, 84.589321E, 25.X.2008, leg. P.B. Budha. CDZMTU065b/16 shells and CDZMTU065P/1 specimen (dissected), Kurintar, Chitwan, 527 m,
27.874143N
, 84.586683, 23.VII.2010, leg. P.B.Budha.
Shell.
Measurements (n = 30): SH 13.0-19.0 mm, SW 5.4-6.3 mm, HA 5.0-6.1 mm, WA 2.7-3.5 mm, Wh 8.5-9.5; approx. 2.7
x
higher than wide, oblong turreted, yellowish brown. Surface with shallow radial ribs, stronger towards the suture, sculpture regular, thin transparent periostracum. Suture deep, spire convex. The first whorl rounded (Fig. 2F1). Aperture small and ovate, 1.7
x
higher than wide, peristome thin, columellar margin short and abruptly truncated.
Genitalia
(n = 2) (Figs 8A). Vas deferens long with a constant diameter. The flagellum is a simple sac. Epiphallic caecum is longer than the flagellum. Epiphallus much shorter than the penis. The penial retractor muscle far apart from the flagellum. Gametolytic duct slender, ending into a balloon-like gametolytic sac, which is slightly longer than the gametolytic duct. Vagina as long as the penis. Interior wall of the penis with a strong fold.
Figure 8. Genitalia of
R. cf. mastersi
and
Rishetia
sp.: A General view of
R. cf. mastersi
, CDZMTU065P B Interior of penis of the same, Kurintar, Chitwan, degraded riverine bushes with big boulders, mixed
Shorea robusta
forest C General view of
Rishetia
sp. CDZMTU078P D Interior of penis of the same, Boshikharka, Dhading, Central Nepal.
Remarks.
A single specimen of
Rishetia
sp. was collected from the
Shorea robusta
forest at Bosikharka, Dhanding (CDZMTU078P). Its shell is similar to
R. mastersi
Godwin-Austen, 1920, but it is slightly smaller and has relatively stronger radial ribs. The shape of its flagellum and epiphallus is similar to that of
R. mastersi
too (Figs 8A, 8C), but the interior of its penis shows three longitudinal folds (Fig. 8D), whereas in
R. mastersi
there is only one longitudinal fold (Fig. 8B).