An integrated taxonomic revision of Ctonoxylon (Coleoptera, Curculionidae, Scolytinae) reveals new Malagasy species originating from multiple recent colonisations of the island Author Jordal, Bjarte H. 0000-0001-6082-443X Department of Natural History, University Museum of Bergen, University of Bergen, P. O. 7800, NO- 5020 Bergen, Norway text ZooKeys 2024 2024-05-28 1203 95 130 journal article 10.3897/zookeys.1203.123757 6FE93D4B-8104-45F4-850A-C1B78CA10A9D Ctonoxylon atrum Browne stat. rev. Figs 68 , 71 , 74 Ctonoxylon atrum Browne, 1965: 190 . Type material. Holotype , male: Nigeria , Idanre [GIS: 7.12, 5.10 ], 30. 9. 1964 , ex Canthium [ NHMUK ]. Diagnosis. Length 3.4 mm . 2.1–2.2 × as long as broad; colour pitch black. Male and female frons slightly impressed and impunctate on lower half, finely granulated above with short scattered setae; eye parts separated by 2 / 3 the size of upper half; anterior lateral margin of prothorax with elongated groove near procoxa; propleural pit present just above procoxa; pronotum with large and deep irregular punctures; elytral suture buckled at midlength, strial punctures longitudinally elongated, subquadrate; interstriae rough, with irregular punctures and rugosities; elytral apex emarginated; interstrial vestiture of confused rows of short, black, scale-like setae. Distribution. Ghana , Cameroon (new country). New record. Cameroon , Limbe, Ekande [GIS: 4.081 , 9.172 ], 19. Nov. 2007 , ex water liana, B. Jordal, leg [1, ZMUB ]. Remarks. This species is rather similar to C. methneri and was synonymised with C. hamatum by Schedl (1970) . However, C. atrum is darker, with shorter elytral setae, the strial punctures are narrowly elongated and more separated, and the pronotal punctures are larger. It is genetically clearly separated from C. methneri and instead forms a sister relationship with C. quadrispinum sp. nov. (Fig. 11 ).