Australian Goblin Spiders Of The Genus Ischnothyreus (Araneae, Oonopidae) Author Edward, Karen L. Author Harvey, Mark S. text Bulletin of the American Museum of Natural History 2014 2014-06-06 2014 389 1 144 http://www.bioone.org/doi/abs/10.1206/865.1 journal article 10.1206/865.1 0003-0090 5380163 Ischnothyreus hamatus , new species Figures 70–71 , map 13 TYPES: AUSTRALIA : Queensland : Male holotype and female allotype from Upper Brookfield , 27.5 ° S , 152.9167 ° E , 100 m ( 28 Jan 1982 , R. Raven , V. Davies ), deposited in QM ( ³ holotype : QM S16002 , PBI _ OON 00005886 ; ♀ allotype : QM S16002 , PBI _ OON 00005596 ) . ETYMOLOGY: The specific epithet is of the Latin hamatus meaning ‘‘hooked’’ ( Brown, 1956 ), and refers to the hooked distal tip of the male palp. DIAGNOSIS: This species can be easily recognized by the curved and pointed distal tip of the male palpal embolic region (fig. 71E), the greatly enlarged base of the male fang, which also exhibits a thin process that curves around the anterior aspect (fig. 71C, D). The female possesses a heavily sclerotized postepigastric scutum and epigynal area. The process overhanging an indistinct epigynal atrium has rectangular-shaped edges and is most heavily sclerotized at the base of the convoluted duct (figs. 70I, 71F). MALE (PBI_OON_005886, figs. 70A–C, G–H, 71A–E). Total length 1.63. CEPHALO- THORAX: Carapace orange-brown, broadly oval in dorsal view, pars cephalica strongly elevated in lateral view, anteriorly narrowed to 0.49 times its maximum width or less, anterolateral corners without extension or projections, surface of elevated portion of pars cephalica smooth, sides strongly reticulate. Clypeus margin unmodified, sinuous in front view, high, ALE separated from edge of carapace by their radius or more; setae light. Eyes: ALE largest, ALE circular, PME circular, PLE oval; posterior eye row straight from above; ALE touching, ALE-PLE touching. Sternum longer than wide, orange-brown, uniform; setae dark, evenly scattered. Chelicerae, endites, and labium pale orange. Chelicerae straight, anterior face unmodified; promargin with two larger denticles; fang dorsoventrally flattened and strongly bent medially, with prominent basal process, anterior aspect with thin curved process (fig. 71C); setae light. Labium elongated hexagon, not fused to sternum, anterior margin indented at middle, same as sternum in sclerotization; without setae. Endites anteromedian tip with one strong, toothlike projection, same as sternum in sclerotization. ABDOMEN: Ovoid; dorsum soft portions white. Book lung covers ovoid. Dorsal scutum orange-brown, covering full length of abdomen, no soft tissue visible from above, middle surface smooth, sides smooth. Epigastric scutum small lateral sclerites absent. Postepigastric scutum orange-brown, covering about 2/3 of abdominal length. Dorsum, epigastric area, and postepigastric area setae dark. LEGS: Pale orange, without color pattern; patella plus tibia I shorter than carapace. Leg spination: femora: I p0-1-1; II p0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Palp proximal segments dark red-brown; embolus dark; femur shorter than trochanter; patella longer than femur; cymbium dark red-brown; bulb dark red-brown, 1 to 1.5 times as long as cymbium, stout; embolic region elongate, obtusely bent, curved strongly toward prolateral aspect, tapered to fine hooked tip, small transparent membranous tissue visible (fig. 71A, B, E). Fig. 70. Ischnothyreus hamatus , sp. nov. Holotype male (PBI_OON 00005886): A. habitus, dorsal view; B. habitus, ventral view; C. habitus, lateral view; G. carapace, anterior view; H. sternum, ventral view. Allotype female (PBI_OON 00005596): D. habitus, dorsal view; E. habitus, ventral view; F. habitus, lateral view; I. epigynum, ventral view. Fig. 71. Ischnothyreus hamatus , sp. nov. Holotype male (PBI_OON 00005886): A. left palp, prolateral view; B. left palp, retrolateral view; C. left chelicerae, anterior view; D. left chelicerae, posterior view; E. left palp, dorsal view. Allotype female (PBI_OON 00005596): F. epigynum, ventral view. Scale lines 5 0.1 mm. Map 13. Map of Queensland showing recorded distribution of Ischnothyreus hamatus ( m ). FEMALE (PBI_OON 5596, figs. 70D–F, I, 71F). Total length 1.75. CEPHALOTHO- RAX: Carapace pars cephalica slightly elevated in lateral view, surface of elevated portion of pars cephalica finely reticulate. Clypeus straight in front view, low, ALE separated from edge of carapace by less than their radius. Chelicerae, endites, and labium orange-brown. ABDOMEN: Dorsal scutum dark orange-brown, covering 1/2 to 3/4 of abdomen, more than 1/2 to most of abdomen width. Epigastric scutum small lateral sclerites present. Postepigastric scutum widely hexagonal, covering about 1/3 of the abdominal length. LEGS: Orange-brown. Leg spination: femora: I p0-1-1; II p0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Ventral view: strong horizontal scleriotization anterior to rounded epigynal atrium, clearly joins posteriorly directed lateral apodemes; heavily sclerotized process overhanging epigynal atrium rectangular, straight to slightly sinuous edge, epigynal atrium shape not distinct but slightly oval; convoluted duct thicker than apodemes (figs. 70I, 71F). OTHER MATERIAL EXAMINED: AUS- TRALIA: New South Wales : Bald Mountain , via Emuvale , rainforest, 1130 m , 28.71667 ° S , 152.26670 ° E , Nov. 17–Dec. 28, 1974 (G. and S. Monteith , QM S16832 , PBI _ OON 5925 ), 3 ³, 2 ♀ ; Bald Mountain , via Emuvale , rainforest, 1130 m , 28.71667 ° S , 152.26670 ° E , Nov. 17–Dec. 28, 1974 (G. and S. Monteith , QM S16839 , PBI _OON 26228), 1 ♀ . Queensland : 0.7 km SW of McAfee’s Lookout , wet sclerophyll forest, 150 m , 27.43333 ° S , 152.86666 ° E , July 6–Oct. 18, 1999 (G. Monteith , QM S78945, PBI _OON 20837), 1 ³, 2 ♀ ; 1 km E of One Tree Hill , semievergreen vine thicket, 180 m , 25.28333 ° S , 151.91666 ° E , Sept. 27–Dec. 14, 1999 (D. and I. Cook , QM S78942, PBI _OON 20840), 1 ³ ; 1 km SW of McAfee’s Lookout , wet sclerophyll forest, 140 m , 27.43583 ° S , 152.86900 ° E , Oct. 18, 1999 , to Feb. 6, 2000 (G. Monteith , QM S57390 , PBI _OON 22101), 4 ³, 4 ♀ ; 3 km SE of ‘‘ Stockhaven ,’’ 450 m , 25.80000 ° S , 151.98333 ° E , Oct. 10–Dec. 19, 1998 (G. Monteith , C. Gough , QM S49538 , PBI _OON 22062), 1 ³ ; 3 km SE of ‘‘ Stockhaven ,’’ 450 m , 25.80050 ° S , 151.98420 ° E , Dec. 19, 1998 , to Jan. 25, 1999 ( G. Monteith , C. Gough , QM S69563, PBI _OON 22162), 1 ³ ; 3 km SE of ‘‘ Stockhaven ,’’ rainforest, 450 m , 25.80000 ° S , 151.98333 ° E , Oct. 10–Dec. 19, 1998 (G. Monteith , C. Gough , QM S79855 , PBI _OON 26265), 1 ³ ; 5.5 km E of One Tree Hill , semievergreen vine thicket, 120 m , 25.33583 ° S , 151.90970 ° E , Sept. 26–Dec. 14, 1999 (D. and I. Cook , QM S72916 , PBI _OON 22006), 1 ³ ; 5.5 km E of One Tree Hill , 120 m , 25.33583 ° S , 151.90970 ° E , Dec. 14, 1999 , to Mar. 19, 2000 (G. and S. Monteith , QM S57855, PBI _OON 22247), 1 ³ ; Bahrs Scrub , Beenleigh , 100 m , 27.75000 ° S , 153.16670 ° E , Dec. 10, 1991 , to Jan. 21, 1992 ( D.J. Cook , QM S25027 , PBI _OON 21724), 1 ³ ; Bare Rock , 2 km N Mount Cordeaux , 1100 m , 28.03333 ° S , 152.38330 ° E , Dec. 31, 1993 , to Feb. 20, 1994 ( G. Monteith , QM S49210 , PBI _OON 22081), 1 ³ ; Beerwah Forestry Reserve (A2), 26.85000 ° S , 152.95000 ° E , June 27, 1990 (M. Glover , QM S33552 , PBI _OON 22017), 1 ³ ; Beerwah Forestry Reserve (A3), 26.85000 ° S , 152.95000 ° E , June 6, 1990 ( Estuarine Research Group , QM S63464 , PBI _OON 21792), 1 ♀ ; Beerwah Forestry Reserve (A3), 26.85000 ° S , 152.95000 ° E , Sept. 19, 1990 (M. Glover , QM S63470, PBI _OON 22013), 1 ³ ; Beerwah Forestry Reserve (A4), heathland, 26.85000 ° S , 152.95000 ° E , June 26, 1991 (M. Glover , QM S69635 , PBI _OON 21795), 1 ³ ; Beerwah Forestry Reserve (A4), 26.85000 ° S , 152.95000 ° E , May 16, 1990 (M. Glover , QM S48814 , PBI _OON 21948), 2 ♀ ; Beerwah Forestry Reserve (A4), heathland, 26.85000 ° S , 152.95000 ° E , June 26, 1991 (M. Glover , QM S63784 , PBI _OON 22104), 1 ³, 1 ♀ ; Beerwah Forestry Reserve (A5), 26.85000 ° S , 152.95000 ° E , June 26, 1991 (M. Glover , QM S63129, PBI _ OON 21971), 1 ♀ ; Beerwah Forestry Reserve (D4), heathland, 26.85000 ° S , 152.95000 ° E , July 18, 1990 (M. Glover , QM S19506 , PBI _OON 21945), 1 ♀ ; Beerwah Forestry Reserve (D4), heathland, 26.85000 ° S , 152.95000 ° E , Aug. 29, 1990 (M. Glover , QM S18937 , PBI _OON 22045), 1 ♀ ; Beerwah Forestry Reserve (E3), heathland, 26.85000 ° S , 152.95000 ° E , Jan. 2, 1991 (M. Glover , QM S19504 , PBI _OON 22015), 1 ♀ ; Beerwah Forestry Reserve (A3), 26.85000 ° S , 152.95000 ° E , Feb. 27, 1991 (M. Glover , QM S63565 , PBI _OON 21942), 1 ³ ; Beerwah Forestry Reserve (A4), 26.85000 ° S , 152.95000 ° E , Aug. 29, 1990 (M. Glover , QM S24002 , PBI _ OON 7246 ), 1 ♀ ; Bellthorpe , 26.83333 ° S , 152.68330 ° E , Mar. 9–May 05, 1997 (G. Monteith , QM S37682 , PBI _OON 21988), 1 ♀ ; Belmont Hills Bushlands , site 1, 27.50784 ° S , 153.11750 ° E , Apr. 22, 2003 (C. Burwell , S. Wright , E. Volschenk , QM S62326 , PBI _OON 22199), 1 ³ ; Black Rock Scrub , rainforest, 350 m , 28.12200 ° S , 152.65800 ° E , Dec. 2, 2000 , to May 13, 2001 ( G. Monteith , QM S57470, PBI _OON 25913), 1 ³ ; Brookfield , Gold Creek Reservoir , 100 m , 27.50000 ° S , 152.91670 ° E , Aug. 19, 1980 (R.J. Raven , G. Gordh , QM S16001 ), 1 ³, 2 ♀ ; Buhot Creek , Burbank , riparian forest, leaf litter, 27.58783 ° S , 153.16980 ° E , Feb. 18, 2004 ( Queensland Museum Party , QM S65763 , PBI _OON 22421), 1 ³ ; Buhot Creek , Burbank , riparian forest, 27.58783 ° S , 153.16980 ° E , July 28–Sept. 2, 2003 ( Queensland Museum Party , QM S65761 PBI _OON 25909), 1 ♀ ; Cameron’s Scrub , Snig track, 120 m , 27.50817 ° S , 152.72380 ° E , Jan. 13– May 16, 1999 (G. Monteith , QM S69587 , PBI _OON 22012), 1 ³ ; Camerons Scrub , snig track, 120 m , 27.50817 ° S , 152.72380 ° E , Nov. 11, 1998 , to Jan. 13, 1999 (G. Monteith , D. Cook , G. Thompson , QM S69607 , PBI _OON 22205), 3 ³ ; Camerons Scrub , snig track, 80 m , 27.51667 ° S , 152.73330 ° E , Jan. 13–May 16, 1999 (G. Monteith , QM S54102 , PBI _OON 22029), 1 ♀ ; Camira , 50 m , 27.63333 ° S , 152.91670 ° E , Feb. 11, 1985 (R.J. Raven , QM S16472 , PBI _OON 26230), 1 ³ ; Cooloola National Park , L Poona , 20 m , 26.20000 ° S , 153.05000 ° E , Oct. 9–19, 1978 (G. Monteith , QM S16036 , PBI _ OON 5909 ), 1 ³ ; Cooloola Village turnoff (CV1), 26.08333 ° S , 153.08330 ° E , July 8– Dec. 4, 1998 (R. Raven , P. Lawless , QM S53391 , PBI _OON 21964), 2 ♀ ; Cooloola Village turnoff (CV1), 26.08333 ° S , 153.08330 ° E , July 8–Dec. 4, 1998 (R. Raven , P. Lawless , QM S51756 , PBI _OON 22014), 3 ♀ ; Cunninghams Gap , 790 m , 28.05000 ° S , 152.40000 ° E , Jan. 6–Mar. 01, 1992 (D.J. Cook , QM S25047 , PBI _OON 21738), 1 ³ ; Cunninghams Gap , 790 m , 28.05000 ° S , 152.40000 ° E , Jan. 6–Mar. 1, 1992 (D.J. Cook , QM S25050 , PBI _OON 21759), 1 ♀ ; D’Aguilar National Park , Mount Glorious Walk , rainforest, leaf litter, 27.32194 ° S , 152.76250 ° E , May 12, 2007 (K. Edward and K. Pitz , QM S96007 , PBI _OON 25732), 1 ³ ; Deer Reserve , via Kilcoy , rainforest, 460 m , 26.95000 ° S , 152.56670 ° E , Mar. 29–June 1, 1975 (G. and S. Monteith , QM S12939 , PBI _ OON 6430 ), 1 ♀ ; Dwyer Creek , Imbil , 100 m , 26.45883 ° S , 152.63480 ° E , May 27, 2002 (G. Monteith , QM S69614 , PBI _OON 22179), 1 ♀ ; Enoggera Reservoir , site 3, 120 m , 27.44067 ° S , 152.91980 ° E , Nov. 10, 1999 (G. Monteith , G. Thompson , QM S57868, PBI _OON 22219), 2 ³, 2 ♀ ; Enoggera Reservoir , site 3, 120 m , 27.44067 ° S , 152.91980 ° E , Nov. 15, 1999 (G. Monteith , QM S57676, PBI _OON 22265), 3 ♀ ; Enoggera Reservoir , site 3, 120 m , 27.44067 ° S , 152.91980 ° E , Jan. 27–Mar. 15, 2000 (G. Monteith , J. Holt , QM S57291 , PBI _OON 22097), 1 ³ ; Enoggera Reservoir , site 4, rainforest, 110 m , 27.44117 ° S , 152.92050 ° E , Aug. 7–Oct. 16, 1999 (G. Monteith , QM S78941, PBI _OON 20856), 1 ³ ; Enoggera Reservoir , site 4, 110 m , 27.44117 ° S , 152.92050 ° E , Mar. 15–18, 2000 (G. Monteith , QM S57365 , PBI _OON 22111), 1 ♀ ; Enoggera Reservoir , site 5, 120 m , 27.44067 ° S , 152.91980 ° E , Oct. 16–21, 1999 (G. Monteith , QM S57309 , PBI _OON 22129), 2 ³ ; Enoggera Reservoir , site 8, rainforest, 100 m , 27.43950 ° S , 152.90230 ° E , Dec. 21, 1999 , to Jan. 27, 2000 ( G. Monteith , J. Holt , QM S72911 , PBI _OON 21992), 1 ♀ ; Ewan Maddock Dam (site D/3), closed forest, 26.80000 ° S , 152.98333 ° E , June 26, 1993 (M. Glover , QM S32192 , PBI _ OON 7331 ), 1 ♀ ; Ewan Maddock Dam (site E/2), heathland, 26.80000 ° S , 152.98330 ° E , Nov. 21, 1993 , to Mar. 22, 1994 (M. Glover , QM S28517 , PBI _OON 21758), 1 ³ ; Fairlies Knob , 0.5 km S, vine forest, 300 m , 25.51666 ° S , 152.28333 ° E , July 21–Oct. 20, 2000 (D. Cook , S. Wright , E. Vanderduys , QM S78702 , PBI _ OON 7146 ), 1 ♀ ; Joalah National Park , rainforest, leaf litter, 380 m , 27.91666 ° S , 153.20000 ° E , Mar. 14, 1973 (R.W. Taylor , ANIC , PBI _OON 25873), 1 ³ ; Kroombit Tops ( Site 8), open forest, leaf litter, 740 m , 24.36666 ° S , 150.95000 ° E , Dec. 11, 1983 (G. Monteith , V. Davies , J. Gallon , G. Thompson , QM S12980 , PBI _OON 26232), 1 ³ ; Lamington IBISCA 0.6 km N of Ballanju Falls , rainforest, leaf litter, 460 m , 28.33333 ° S , 153.33333 ° E , Mar. 19, 2008 (S. Wright , A. Nakamura , QM S86459, S16111 ), 2 ♀ ; Lamington National Park , O’Reilly’s , rainforest, leaf litter, 28.23333 ° S , 153.13333 ° E , Nov. 22–27, 1978 ( Lawrence , Weir , ANIC , PBI _ OON 5571 ), 1 ♀ ; Lamington National Park , O’Reilly’s , rainforest, 28.23333 ° S , 153.13333 ° E , Nov. 1, 1980 , to Jan. 30, 1981 (J. Grimshaw , QM S78253, PBI _ OON 7227 ), 1 ♀ ; Lamington Np IBISCA, 0.5 km SSE Binna Burra Lodge , rainforest, leaf litter, 770 m , 28.33333 ° S , 153.31666 ° E , Mar. 18, 2008 (S. Wright , A. Nakamura , QM S86438 , S86448, S86449 ), 1 ³, 4 ♀ ; Lamington National Park IBISCA, 300 B, rainforest, leaf litter, 300 m , 28.00000 ° S , 153.00000 ° E , Jan. 25, 2008 (S. Wright , A. Nakamura , QM S86453 , PBI _OON 26267), 1 ♀ ; Lamington National Park IBISCA, 500 A, rainforest, leaf litter, 560 m , 28.35000 ° S , 153.23333 ° E , Jan. 28, 2008 (S. Wright , A. Nakamura , QM S86454 ), 1 ♀ ; Lamington National Park IBISCA, 500 B, rainforest, 28.35000 ° S , 153.23333 ° E , Jan. 15–24, 2007 (K. Staunton , QM S86463, PBI _OON 26264), 1 ³ ; Lamington National Park IBISCA, 500 C, rainforest, leaf litter, 500 m , 28.00000 ° S , 153.00000 ° E , Jan. 28, 2008 (S. Wright , A. Nakamura , QM S86440, S86447 , S86460 , S86461, S86456 ), 2 ³, 4 ♀ ; Lamington National Park IBISCA, 700B, rainforest, leaf litter, 775 m , 28.31666 ° S , 153.20000 ° E , Jan. 18, 2008 (S. Wright , QM S86458 , PBI _OON 26263), 1 ³, 1 ♀ ; Lamington National Park IBISCA, 900 A, rainforest, 900 m , 28.00000 ° S , 153.00000 ° E , Jan. 15–24, 2007 (K. Staunton , QM S78045 , S86450, S86451 ), 1 ³, 2 ♀ ; Lamington National Park , Binna Burra , leaf mold, rotten wood, 28.18333 ° S , 153.18333 ° E , Sept. 7, 1952 (T.E. Woodward , QM S78044 , PBI _OON 26249), 1 ♀ ; Lamington National Park , IBISCA Plot # IQ–300–A, rainforest, leaf litter, 267 m , 28.14777 ° S , 153.13694 ° E , Jan. 25, 2008 (S. Wright , A. Nakamura , QM S86439 , S86440, S86445 , S86462 , S86464 ), 3 ³, 3 ♀ ; Lower Coomera , 350 m , 28.18333 ° S , 153.18330 ° E , Dec. 3, 1994 , to Jan. 9, 1995 (G. Monteith , H. Janetzki , QM S31904 , PBI _OON 22263), 4 ³, 2 ♀ ; Lower Coomera , 350 m , 28.18333 ° S , 153.18330 ° E , Jan. 9–Apr. 6, 1995 (G. Monteith , QM S49208 , PBI _OON 22420), 1 ³, 1 ♀ ; Malaan SF, rainforest, leaf litter, 1000 m , 17.58333 ° S , 145.58330 ° E , Apr. 20–24, 1978 (R. Raven , V. Davies , QM S16126 , PBI _OON 25916), 1 ³ ; Miala National Park , rainforest, leaf litter, 635 m , 27.20000 ° S , 152.46000 ° E , Mar. 13, 1973 (R.W. Taylor , ANIC , PBI _OON 25849), 1 ³, 1 ♀ ; Mistake Mountains (middle), 950 m , 27.96667 ° S , 152.38330 ° E , Jan. 9–Feb. 13, 1977 (G. and S. Monteith , QM S16823 , PBI _OON 25956), 2 ♀ ; Mount Coot-tha , Brisbane , leaf litter, 27.48334 ° S , 152.95000 ° E , Feb. 1, 1980 (R. Raven , V. Davies , QM S16175 , PBI _ OON 5924 ), 1 ♀ ; Mount Coot-tha , Brisbane , 27.48334 ° S , 152.95000 ° E , June 6, 1980 (R. Raven , V. Davies , QM S16109 , PBI _OON 25886), 1 ³, 1 ♀ ; Mount Coot-tha , Brisbane , leaf litter, 27.48334 ° S , 152.95000 ° E , Feb. 20, 1980 (R.J. Raven , QM S16181 , PBI _OON 26227), 1 ♀ ; Mount Cotton , 200 m , 27.60967 ° S , 153.20350 ° E , Sept. 3–Dec. 12, 1997 (G. Monteith , QM S60473 , PBI _OON 22090), 2 ³, 2 ♀ ; Mount Cotton , upper gully, 150 m , 27.60967 ° S , 153.20530 ° E , Dec. 12, 1997 , to May 7, 1998 (G. Monteith , QM S60085, PBI _OON 22136), 2 ³, 2 ♀ ; Mount Glorious , 690 m , 27.33333 ° S , 152.76670 ° E , Sept. 20, 1979 (G. Monteith , QM S16049 , PBI _ OON 5905 ), 1 ♀ ; Mount Glorious , 760 m , 27.33333 ° S , 152.76666 ° E , May 1, 1970 (K. Plowman , ANIC S16164 , PBI _ OON 5915 ), 2 ♀ ; Mount Glorious , 760 m , 27.33333 ° S , 152.76666 ° E , Oct. 15, 1970 (H. Williams , ANIC S16184 , PBI _ OON 5928 ), 1 ♀ ; Mount Glorious , wet sclerophyll forest, leaf litter, 760 m , 27.33333 ° S , 152.76666 ° E , Sept. 18, 1968 (G. Monteith , ANIC , PBI _OON 25852), 1 ³, 1 ♀ ; Mount Glorious , 690 m , 27.33333 ° S , 152.76670 ° E , June 7–20, 1974 (V.E. Davies , QM S16193 , PBI _OON 26234), 1 ³, 1 ♀ ; Mount Goonaneman , 670 m , 25.43333 ° S , 152.13330 ° E , Nov. 3–7, 1980 (V. Davies , R. Raven , QM S16149 , PBI _ OON 5918 ), 9 ³, 15 ♀ ; Mount Mee , Neurum Creek , 210 m , 27.08333 ° S , 152.70000 ° E , Oct. 28, 1977 , to Jan. 20, 1978 ( G. and S. Monteith , QM S12988 , PBI _ OON 5916 ), 2 ♀ ; Mount Superbus , base, rainforest, 1350 m , 28.23333 ° S , 152.48333 ° E , Mar. 12–June 13, 1990 (T.B. Churchill , R.J. Raven , QM S15884 , PBI _OON 26231), 1 ♀ ; Mount Superbus , half ascent, 28.73333 ° S , 152.48330 ° E , June 13–Oct. 30, 1990 (T. Churchill , R. Raven , K. Williams , QM S26287 , PBI _OON 21995), 1 ³, 2 ♀ ; Mount Superbus , halfway. Site 3, 28.23333 ° S , 152.48330 ° E , Oct. 30, 1990 (T. Churchill , R. Raven , K. Williams , QM S25890 , PBI _OON 21979), 1 ♀ ; Mount Tamborine National Park , rainforest, 27.88333 ° S , 153.18333 ° E , Feb. 21–Mar. 29, 1984 (J. Grimshaw , QM S78260, PBI _ OON 7228 ), 1 ³ ; Mount Tamborine National Park , Palm Grove , rainforest, 27.88333 ° S , 153.18333 ° E , Oct. 13, 1956 (T.E. Woodward , QM S78049 , PBI _OON 26248), 1 ♀ ; Mount Tenison Woods , 620 m , 27.32333 ° S , 152.72170 ° E , May 15, 1997 (G. Monteith , QM S43063 , PBI _OON 21976), 1 ♀ ; Mount Tenison Woods , 760 m , 27.30000 ° S , 152.75000 ° E , Mar. 22, 1979 (G. Monteith , QM S16071 , PBI _OON 26226), 1 ³ ; Mudlo gap, bottom, rainforest, 170 m , 26.01666 ° S , 152.23333 ° E , Oct. 9–Dec. 17, 1998 (G. Monteith , C. Gough , QM S78706, PBI _ OON 7144 ), 1 ³ ; Samford , dry eucalypt woodland, 320 m , 27.36666 ° S , 152.88333 ° E , Sept. 18, 1968 (G. Monteith , ANIC , PBI _OON 25858), 1 ³ ; Searys Scrub , Cooloola National Park , 26.20000 ° S , 153.05000 ° E , Feb. 3–8, 1976 (R. Raven , V. Davies , QM S16147 , PBI _ OON 5929 ) 10 ³, 10 ♀ ; Tamborine Mountain , Joalah National Park , rainforest, leaf litter, 27.93333 ° S , 153.20000 ° E , Oct. 18–21, 1978 ( Lawrence , Weir , ANIC , PBI _ OON 5569 ), 1 ³ ; Tamborine Mountain , Joalah National Park , rainforest, leaf litter, 27.93333 ° S , 153.20000 ° E , Oct. 18–21, 1978 ( Lawrence , Weir , ANIC , PBI _ OON 5570 ), 1 ♀ ; Tamborine National Park , Curtis Falls Track , rainforest, leaf litter, 27.93194 ° S , 153.19388 ° E , May 11, 2007 (K. Edward and K. Pitz , QM S96008 , PBI _OON 25731), 1 ³ ; Tamborine National Park , Curtis Falls Track , rainforest, leaf litter, 27.93194 ° S , 153.19388 ° E , May 11, 2007 (K. Edward and K. Pitz , QM S96009 , PBI _OON 25733), 2 ³, 1 ♀ ; Tullawallal , Binna Burra , 950 m , 28.20000 ° S , 153.20000 ° E , Dec. 3, 1994 , to Jan. 9, 1995 ( G. Monteith , H. Janetzki , QM S37823 , PBI _OON 22116), 1 ³ ; Upper Brookfield , 100 m , 27.50000 ° S , 152.91670 ° E , June 2, 1981 (V. Davies , R. Raven , QM S16476 , PBI _ OON 5911 ), 1 ♀ ; Upper Brookfield , 100 m , 27.50000 ° S , 152.91670 ° E , Nov. 14– 28, 1980 (V. Davies , R. Raven , QM S16480 , PBI _OON 25953), 1 ³ ; Upper Brookfield , 100 m , 27.50000 ° S , 152.91670 ° E , July 3, 1981 (V. Davies , R. Raven , QM S16495 , PBI _OON 25954), 1 ♀ ; Upper Brookfield , 100 m , 27.50000 ° S , 152.91670 ° E , Apr. 8–23, 1981 (V. Davies , R. Raven , QM S16697 , PBI _OON 25955), 3 ♀ ; Upper Brookfield , 100 m , 27.50000 ° S , 152.91670 ° E , Jan. 28, 1982 (V. Davies , R. Raven , QM S16002 , PBI _OON 25958), 4 ³, 1 ♀ ; Upper Brookfield , 100 m , 27.50000 ° S , 152.91670 ° E , June 17, 1981 (V. Davies , R. Raven , QM S16492 , PBI _OON 26229), 2 ³ ; Upper Brookfield , 100 m , 27.50000 ° S , 152.91670 ° E , Aug. 14, 1981 (V. Davies , R. Raven , QM S16491 , PBI _OON 26233), 2 ♀ ; Upper Brookfield , 100 m , 27.50000 ° S , 152.91670 ° E , Nov. 14–28, 1980 (V. Davies , R. Raven , QM S16694 , PBI _OON 26235), 1 ♀ ; Upper Brookfield , 100 m , 27.50000 ° S , 152.91670 ° E , Jan. 12, 1982 (R.J. Raven , QM S16000 , PBI _OON 26237), 2 ♀ ; Upper Brookfield , 100 m , 27.50000 ° S , 152.91670 ° E , Mar. 19, 1982 (R.J. Raven , QM S12998 PBI _ OON 5923 ), 2 ♀ . DISTRIBUTION: This species is relatively widespread from Bald Mountain near the border of New South Wales to north of Bundaberg, in northeastern Queensland (map 13).