Revision of the funnel-web spider genus Novalena (Araneae: Agelenidae) Author Maya-Morales, Julieta Author Jiménez, María Luisa text Zootaxa 2017 4262 1 1 88 journal article 33105 10.5281/zenodo.571683 5ab1c651-825a-43bc-af00-11b77a792de3 1175-5326 571683 894795FB-ED38-4132-92B9-6AC8C3E41566 Novalena annamae (Gertsch & Davis 1940) Figure 2 ; Map 3 Agelena annamae Gertsch & Davis 1940: 2 , figures 3, 7–8 ( ♂ ♀ ) Novalena annamae Chamberlin & Ivie 1942 : 229 Type . Holotype female : MEXICO : Michoacán : Municipality of Zitácuaro , Zitácuaro , 19°21′N , 100°28′W , 27.VI.1936 , L.I. Davis , deposited at AMNH (examined). Diagnosis . The males of N. annamae differ from all other species by a membranous fulcrum near the base of the embolus ( Fig. 2 a) (absent in all other species). The females are similar to N. mexiquensis in having the atrium with lateral sclerotized tips ( Fig. 2 c), but differ from that species in having the atrial spurs separated from posterior margin by three times their length ( Fig. 2 c). Description . Male ( allotype ): Carapace light brown. Chelicerae brown, condyles light brown. Endites and labium light brown with white tips. Sternum with two longitudinal black bands. Legs light brown with three rings on femur. Opisthosoma light brown with an anterior dark brown spot, two lateral bands, and three posterior chevrons. Spinnerets with ALS light brown, PLS with dark brown basal part and light brown distal part. Total length 7.5. Carapace length 3.8, width 2.4, cephalic region width 1.3, ocular region width 0.8. Chelicerae with three retromarginal teeth; basal segment length 1.4, fang length 0.7. Labium wider than long (0.5/0.4). Endites slightly convergent (distance at their base compared at their tips 0.5/0.3). Sternum longer than wide (1.8/1.43). Opisthosoma longer than wide (4/2). ALS separated by their basal diameter (0.3/0.3), PLS with distal segment slightly longer than basal segment (0.5/0.4). Leg lengths: I—femur 3.2/ patella-tibia 4.05/ metatarsus 3.2/ tarsus 2.3; II—2.8/ 3.6/ 3/ 2; III—2.6/ 3.4/ 3.2/ 1.8; IV—3.8/ 5.72/ 4.85/ 2.55. Spination: femur dorsal I—1-2 -3/ II—1-3 - 3/ III—1-3 -3/ IV—1-2 -3; patella I—dorsal 1-1/ prolateral 1/ retrolateral 0; II—1-1 / 1/ 0; III—1-1 / 1/ 1; IV—1-1 / 1/ 1; tibia I—dorsal 0/ ventral 2-2-2/ prolateral 1-1-0/ retrolateral 0; II—0/ 2-2-2/ 1-1-0/ 0; III—1-1 -0/ 2-2-2/ 1-1-0/ 1- 1-0; IV—1-1 -0/ 2-2-2/ 1-1-0/ 1-1-0; metatarsus I—0/ 2-2-2/ 0-1-1/ 0-1-1; II—0-0-2/ 2-2-2/ 0-1-1/ 0-1-1; III—1-1 - 2/ 2-2-2/ 1-1-1/ 1-1-1; IV—1-2 -2/ 2-1-2-2/ 1-1-1/ 1-1-1. Number of trichobothria: I—6 , II—6 , III—6 , IV—6. Number of dorsal spines on pedipalp: femur 3, tibia?. Cymbium length 1.2, width 0.8. Embolus curved; conductor with ventral projection slightly larger than dorsal projection; tegular median process with one rounded projection; fulcrum present; dorsal projection of RTA with two subprocesses, ventral projection slightly flattened ( Fig. 2 a, b). FIGURE 2 . Novalena annamae (Gertsch & Davis 1940) : a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; CD, copulatory ducts; CO, copulatory openings; E, embolus; F, fulcrum; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S1, primary spermathecae; S2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 0.5 mm; e = 0.25 mm. Female ( holotype ): Coloration similar to male, slightly darker. Total length 9.6. Carapace length 4.45, width 2.8, cephalic region width 1.6, ocular region width 1. Chelicerae with three retromarginal teeth; basal segment length 1.9. Labium wider than long (0.63/0.5). Endites convergent (distance at their base compared at their tips 0.63/0.3). Sternum longer than wide (2.1/1.65). Opisthosoma longer than wide (5.4/3.15). ALS separated by more their basal diameter (0.4/0.3), PLS with distal segment slightly longer than basal segment (0.6/0.5). Leg lengths: I—femur 3.25/ patella-tibia 4.3/ metatarsus 3/ tarsus 2.05; II—3.1/ 3.7/ 2.8/ 1.8; III—3.1/ 3.6/ 2.9/ 1.7; IV—4.1/ 1.5/ 3.45/ 2.25. Spination: femur dorsal I—1-1 -3/ II—1-3 -3/ III—1-3 -3/ IV—1-1 -3; patella I—dorsal 1-1/ prolateral 1/ retrolateral 0; II—1-1 / 1/ 0; III—1-1 / 1/ 1; IV—1-1 / 1/ 1; tibia I—dorsal 0/ ventral 2-2-2/ prolateral 1- 1-0/ retrolateral 0; II—0/ 2-2-2/ 1-1-0/ 0; III—1-1 -0/ 2-2-2/ 1-1-0/ 1-1-0; IV—1-1 -0/ 2-2-2/ 1-1-0/ 1-1-0; metatarsus I—0/ 2-2-2/ 1-1-0/ 0; II—0/ 2-2-2/ 1-1-0/ 0; III—2-2 -2/ 2-2-2/ 0-1-1/ 0-1-0; IV—1-1 -2/ 2-1-1-2/ 1-1-1/ 1-1-1. Number of trichobothria on tarsus: I—8 , II—8 , III—8 , IV—8. Dorsal spines on pedipalp femur: 3. Epigynal plate wider than long (1/0.7); atrium in shallow depression, anterior margin with two sclerotized tips; anterior hyaline spurs separated by more their basal width ( Fig. 2 c, e). Internal genitalia: copulatory ducts straight, primary spermathecae adjacent and with a dorsal transverse mark ( Fig. 2 d). Variation . Males (n=5): body length 5.5–7.75; carapace length 3.13–4; patella-tibia I length 3.75–4.75. Females (n=5): body length 7.5–11.38; carapace length 3.88–5; patella-tibia I length 3.88–4.5. Additional material examined . Allotype : MEXICO : same data as holotype, 1 ♂ ( AMNH ). Other material : MEXICO: Michoacán : Municipality of Tancítaro , Apo , 1067 m , 6–10.VI.1942 , H. Wagner , 1 ♂ 1 ♀ ( AMNH ) ; Municipality of Uruapan, Parque Nacional Uruapan, 26.VI.1988 , M.L. García, 1 ♂ 1 ♀ (CAFBUM_AGE001); same data, 1 ♂ (CARCIB) ; same data, 1 ♀ (CARCIB) ; same data except 30.VII.1988 , 1 ♀ (CARCIB). Nuevo León : Municipality of Sabinas Hidalgo , Sabinas Hidalgo , 1 ♂ ( AMNH ) . Distribution . MEXICO : Michoacán , Nuevo León (Map 3).