Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae) Author Likhitrakarn, Natdanai Author Golovatch, Sergei I. Author Prateepasen, Rujiporn Author Panha, Somsak text ZooKeys 2010 72 23 68 http://dx.doi.org/10.3897/zookeys.72.744 journal article http://dx.doi.org/10.3897/zookeys.72.744 1313-2970-72-23 Tylopus extremus sp. n. Figs 79 Holotype ♂ (CUMZ), Thailand, Chiang Mai Province, Fang District, Doi Phahom Pok National Park, 6.07.2009, leg. A. Pansook. Paratypes: 1 ♂, 1 ♀ (CUMZ), 1 ♂ (ZMUC), 1 ♂ (ZMUM), same locality, together with holotype. Name: To emphasize the extremely long spines h and m of the gonopod. Diagnosis: Differs from congeners in process h being subflagelliform while process m extremely long and prominent. Description: Length ca 30 mm (holotype), 27-30 mm (♂), 32.5 mm (♀), width of midbody pro- and metazona 2.0 and 2.9 mm (holotype), 1.9-2.4 and 2.8-3.3 mm (♂), 2.5 and 3.0 mm (♀), respectively. Coloration of live animals, as well as of alcohol material black-brown (Fig. 7 A-G ): calluses of paraterga and antennae only slightly lighter , light brown to brown, venter and legs contrastingly light, yellow (Fig. 7 A-G ), tip of antennae pallid. All characters as in Tylopus bispinosus sp. n., except as follows. Antennae rather short and slender, reaching behind to end of segment 3 (♂) dorsally. Collum with paraterga like rudimentary flaps, especially poorly developed in ♀. In width, head <collum = segments 3-4 <2 <5-16 (♂) (Fig. 7B), or head <collum <segment 3 <2 and 4 <5-16(♀); thereafter body gradually and gently tapering towards telson. Metaterga with two transverse rows of rather long setae: 2+2 in anterior and 2(3)+2(3) in posterior row, the latter often abraded, but then readily traceable as insertion points on low longitudinal ridges or tubercles (Fig. 7 B-G ). Axial line thin, visible on both halves of metaterga. Paraterga strongly developed (Fig. 7 A-G ), lying rather low (at 1/2-1/3 midbody height), slightly inclined laterally, pointed caudally and acutangular already from segment 2, especially strongly so on caudal segments, very clearly surpassing rear contour only on segments 16-19; anterior 1/3 of poreless calluses with two barely visible, lateral, setigerous incisions, but with only a single, likewise poorly developed incision anteriorly on pore-bearing calluses (Fig. 7 B-F ); paraterga slightly less strongly developed in ♀. Transverse sulcus evident on metaterga 5-18, reaching bases of paraterga, evident and rather deep, faintly rugulose at bottom. Stricture between pro- and metazona weakly striolate (Fig. 7 B-G ). Epiproct emarginate at tip, pre-apical papillae evident (Fig. 7G). Hypoproct subtrapeziform, caudal setae widely separated (Fig. 7H). Pleurosternal carinae as compete ridges with a caudal tooth on segments 2-4 (♂) or 2 and 3 (♀), like separated anterior bulges and increasingly poorly developed caudal denticles until segment 16 (Fig. 7C, E, F). Sterna moderately setose, without modifications; an entire, linguiform, sternal lobe between ♂ coxae 4 (Fig. 7I, J). Legs long, in ♂ very distinctly incrassate, 1.7-2.0 or ca 1.3 times as long as midbody height in ♂ and ♀, respectively (Fig. 7B, C, H), ♂ prefemora very distinctly bulged laterally and clothed with dense and adpressed pilosity ventrally (Fig. 7K), acropodites also with similarly dense and ventrally adpressed pilosity, including tarsal brushes. All ♂ postfemora and tibiae except for a few posteriormost ones with a small, but evident adenostyle (= tubercle) at midway on ventral side (Fig. 7K). Gonopods (Figs 8, 9) with lobe l well-demarcated, but unusually prominent, high and elongated; spine h long, extremely slender and subflagelliform; spine z rather short and simple; spine m very prominent, straight and long. Figure 7. Tylopus extremus sp. n., ♂ paratype ( A-K ). A habitus, live coloration B, C anterior part of body, dorsal and lateral views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F, G, H posterior part of body, lateral, dorsal and ventral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively K midbody leg. Figure 8. Tylopus extremus sp. n., ♂ paratype. A, B right gonopod, mesal and lateral views, respectively C-F distal part of right gonopod, mesal, lateral, subcaudal and suboral views, respectively. Scale bar: 0.2 mm. Figure 9. Tylopus extremus sp. n., ♂ paratype. A, B right gonopod, lateral and mesal views, respectively. Scale bar: 0.5 mm.