New gammaroid family, genera and species from subterranean waters of Japan, and their phylogenetic relationships (Crustacea: Amphipoda)
Author
Tomikawa, Ko
Author
Kobayashi, Norio
Author
Morino, Hiroshi
Author
Mawatari, Shunsuke F.
text
Zoological Journal of the Linnean Society
2007
2007-04-30
149
4
643
670
https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2007.00277.x
journal article
5394
10.1111/j.1096-3642.2007.00277.x
7ade6912-d3c9-4d41-8292-ef315e673c9d
0024-4082
4634851
E58C6912-18DC-4D8A-903D-E9EAC01A79FA
LUCIOBLIVIIDAE
TOMIKAWA
FAM. NOV.
Diagnosis
: Mandible: molar weak, not triturative, without seta. Maxilla 1: outer plate with 7 robust serrate setae apically. Maxilliped: outer plate very short, not reaching end of palp article 1. Pereopod 7: basis not expanded. Gills: coxal gills on pereopods 2–6. Uropod 1: peduncle with 3–4 robust basofacial setae. Uropod 3: inner ramus as long as outer ramus; outer ramus elongate, uniarticulate.
Type
genus
:
Lucioblivio
Tomikawa
gen. nov.
Figure 8.
Octopupilla felix
gen. nov., sp. nov.
, holotype, female, 7.7 mm, NSMT-Cr 16655. A, upper lip; B, lower lip; C, left mandible; D, lacinia and incisor of left mandible; E, right mandible; F, lacinia and incisor of right mandible; G, right maxilla 1; H, inner plate of right maxilla 1; I, apical setae on palp of right maxilla 1; J, apical setae on palp of left maxilla 1; K, robust apical setae on outer plate of right maxilla 1; L, maxilla 2; M, maxilliped; N, inner plate of maxilliped; O, outer plate of maxilliped. Scale bars = 0.1 mm.
Figure 9.
Octopupilla felix
gen. nov., sp. nov.
, holotype, female, 7.7 mm, NSMT-Cr 16655. A, gnathopod 1; B, palmar margin of propodus of gnathopod 1 (some setae are omitted); C, dactylus of gnathopod 1; D, gnathopod 2; E, palmar margin of propodus of gnathopod 2 (some setae are omitted); F, dactylus of gnathopod 2. Scale bars = 0.1 mm.
Remarks
:
Luciobliviidae
fam. nov.
can be differentiated from all other gammaroids by the following combination of diagnostic characters: the non-triturative nature of the mandibular molar, the small size of the outer plate of the maxilliped, the lack of coxal gill 7, and the uniarticulate outer ramus of uropod 3.
Luciobliviidae
is similar to
Gammaroporeiidae
Bousfield, 1979
and
Mesogammaridae
Bousfield,
1977
in lacking coxal gill 7. However, from both
Gammaroporeiidae
and
Mesogammaridae
,
Luciobliviidae
is excluded because of the non-triturative nature of the mandibular molar and the small size of the outer plate of the maxilliped. Additionally, it is excluded from
Gammaroporeiidae
by the non-expanded basis of pereopod 7, the presence of basofacial armature on the peduncle of uropod 1, and the elongate outer ramus of uropod 3. Differentiating characters of
Luciobliviidae
,
Gammaroporeiidae
and
Mesogammaridae
are listed in
Table 2
.
Luciobliviidae
has many unusual features. In addition to our previous listing of diagnostic characters which easily separate
Luciobliviidae
from other gammaroids, it may help further studies of this most remarkable family if we list here those features: the lack of marginal setae of the upper lip; the presence of minor prickle-like setae on the tip of the palp article 3 of the mandible; the non-bifid lacinia mobilis of the right mandible; the lack of significant pubescence on the maxillae; the poorly armed inner plate of maxilla 1, with only two plumose setae; the inner plate of maxilla 2 with two plumose facial setae in a oblique row; the poorly armed inner plate of the maxilliped, without a row of inner marginal plumose setae; the presence of small triangular protuberances on the palmar margin of the propodus of the gnathopods; and the lack of associated setae of the retinacula of the pleopods.
At present this family contains only one genus,
Lucioblivio
gen. nov.
from
Japan
.