Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae) Author De Mattia, Willy https://orcid.org/0000-0002-0056-467X Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria willy.demattia@icgeb.org Author Reier, Susanne Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria Author Haring, Elisabeth Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria text ZooKeys 2021 2021-12-14 1077 1 175 http://dx.doi.org/10.3897/zookeys.1077.67081 journal article http://dx.doi.org/10.3897/zookeys.1077.67081 1313-2970-1077-1 C28AD65A76F242CFBED7DFB3702CABCE 734088641608531C8E2CC69397B000ED Charpentieria itala trepida ( Kaeufel , 1928) Figs 48.5, 48.6, 50.4 Distribution: The taxon is known for several stations around the Idro Lake, at the boundary between the provinces of Brescia and Trento ( Nardi 2011 : 98). Examined material: Forte Cima Ora , Anfo , Brescia , Lombardia , Italy . 1500 m asl , 45°47'50.57"N , 10°28'3.60"E , W. De Mattia and J. Macor leg. and det., 2 dissected spm . External morphology of the genital organs (Fig. 48 .5): The V is 3 x longer than the FO. The FDBC is 2 x longer than the SDBC+BC. The BC+SDBC is markedly club-like, short, with clear distinction between the SDBC and the BC. The D is slightly longer than the SDBC+BC and thinner in diameter. The V is cylindrical and long, slightly swollen in its mid part. The PC is ~ 2 x as long as the V. The P is distally pedunculate and abruptly swollen and widens in diameter along its proximal part. The transition between P and EP is not clearly visible. The PR is long and robust. The E is much shorter than the P but wide in diameter. Internal morphology of the genital organs (Fig. 48 .6): The A is smooth. The V presents smooth longitudinal pleats. The P is completely smooth. The PP is folded and elongated. It occupies half of the internal penial volume. The ER is missing. The PP is directly connected with the ELP. The epiphallar formula is: PP(ELP). The E shows two moderately fringed ELP. They proximally fade before the VD. Remarks: In the surroundings of Forte Cima Ora (Anfo) we collected this costate form that was wrongly identified as Charpentieria itala tiesenhauseni (Gredler, 1855) by Nardi (2011 : 98, Fig. 6 ). Charpentieria itala tiesenhauseni seems to be restricted to the type locality (Bocca di Valle) and few additional localities E of the Lago d'Idro (Nordsieck pers. comm.). The population from Forte Cima Ora is here considered as a costate form of C. itala trepida (see Nordsieck 2007 : 54).