Three new species of the extant genus Eucinetus from mid-Cretaceous amber of northern Myanmar (Coleoptera: Eucinetidae) Author Li, Yan-Da 0000-0002-9439-202X State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China & Bristol Palaeobiology Group, School of Earth Sciences, University of Bristol, Bristol BS 8 1 TQ, UK ydli@pku.edu.cn Author Engel, Michael S. 0000-0003-3067-077X Division of Invertebrate Zoology, American Museum of Natural History, New York, NY 10024 - 5192, USA & Facultad de Ciencias Biológicas, Universidad Nacional Mayor de San Marcos, Lima 15081, Perú & Departamento de Entomología, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Lima 15072, Perú mengel@amnh.org Author Huang, Di-Ying 0000-0002-5637-4867 State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China dyhuang@nigpas.ac.cn Author Cai, Chen-Yang 0000-0002-9283-8323 State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China cycai@nigpas.ac.cn text Zootaxa 2024 2024-08-05 5492 2 214 230 http://zoobank.org/880aec67-7542-4e96-98cd-86a2c2a31da3 journal article 10.11646/zootaxa.5492.2.4 1175-5326 13219234 880AEC67-7542-4E96-98CD-86A2C2A31DA3 Eucinetus panghongae Li & Cai sp. nov. ( Figs 5–8 ) Material. Holotype , NIGP203940 , male. Etymology. The species is named after the coleopterist Dr. Hong Pang. Locality and horizon. Amber mine located near Noije Bum Village, Tanai Township, Myitkyina District, Kachin State , Myanmar ; unnamed horizon, mid-Cretaceous, Upper Albian to Lower Cenomanian. Diagnosis. Labrum emarginate ( Fig. 7A ). Apical maxillary palpomere asymmetrical, apically with aciculate tip ( Fig. 7A ). Antennomere 3 about 0.85 times as long as 4 ( Fig. 7A ); antennomere 11 about 2.5 times as long as wide ( Fig. 7B ). Scutellar shield rounded at apex ( Fig. 7F ). Elytra about 1.8 times as long as combined width. Hind wings present. Metanepisternum at least twice as long as wide. Meso- and metatibiae at most with only one or two spines along outer edge ( Figs 6B , 7E ). Mesotibial spurs large; both distinctly stouter than spines along apical fringe of mesotibia ( Fig. 7D ). Metatibial spurs equal ( Fig. 7F ). FIGURE 5. General habitus of Eucinetus panghongae Li & Cai sp. nov. , holotype, NIGP203940, under incident light. A , Dorsal view. B , Ventral view. Scale bars: 500 μm. FIGURE 6. General habitus of Eucinetus panghongae Li & Cai sp. nov. , holotype, NIGP203940, under confocal microscopy. A , Dorsal view. B , Ventral view. Scale bars: 500 μm. Description. Body fusiform and slender, about 1.6 mm long, 0.7 mm wide; surface with fine short setae. Head deflexed against ventral side of prothorax; entirely concealed from above by pronotum. Compound eyes weakly emarginate anteriorly, with interfacetal setae. Antennae 11-segmented, filiform; antennomere 2 elongate; antennomeres 3–10 moderately longer than wide; antennomere 3 about 0.85 times as long as 4; antennomere 11 about 2.5 times as long as wide. Frontoclypeal suture straight anteriorly and with distinct anterolateral angles. Labrum weakly narrowed toward apex, apically emarginate. Mouthparts non-suctorial. Apical maxillary palpomere asymmetrical, apically with aciculate tip. Prothorax widest at base. Lateral pronotal carinae complete, simple. Pronotal disc simple. Procoxae obliquely oriented, well-projecting, narrowly separated. Scutellar shield rounded at apex. Elytra about 1.8 times as long as combined width; transversely strigulate; sutural stria present; epipleuron narrow, incomplete. Hind wings present. Metaventral posterior process with distinct median discrimen. Metanepisternum at least twice as long as wide. Metacoxae contiguous; metacoxal plates well-developed and greatly enlarged. FIGURE 7. Details of Eucinetus panghongae Li & Cai sp. nov. , holotype, NIGP203940, under confocal microscopy. A , Head, dorsal view. B , Antennae. C , Mid leg. D , Fore and mid legs. E , Mid leg. F , Abdominal apex, ventral view. G , Hind leg. H , Scutellar shield, dorsal view. I , Abdominal apex, dorsal view.Abbreviations: a2–11, antennomeres 2–11; cl, clypeus; lb, labrum; mstb, mesotibia; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pts, protarsus; sc, scutellar shield; v5–6, ventrites 5–6. Scale bars: 100 μm. Mid and hind legs with fringe of spines at apices of tibiae and tarsomeres; fore legs without apical fringes of spines. Meso- and metatibiae at most with only one or two spines along outer edge. Mesotibial spurs large; both distinctly stouter than spines along apical fringe of mesotibia. Metatibial spurs equal. Tarsi 5-5-5, simple. Meso- and metapretarsal claws with accessory lobes ( Fig. 7G ). Abdomen with six ventrites. Posterior margin of ventrite 5 subtruncate, not emarginate. Remarks. The sternite VIII of NIGP203940 (and also NIGP203939) is densely covered with setae like the preceding ventrites ( Fig. 7F ). It should be normally exposed when alive, representing the sixth ventrite. In some recent comprehensive coleopteran compendia, the abdomen is described as having only five ventrites for the whole Eucinetidae ( Lawrence & Ślipiński 2013 ; Leschen 2016 ). However, the sternite VIII is actually commonly exposed in many extant members of Eucinetus ( Vit 1977a ) .