Systematics of Sparganothoides Lambert and Powell, 1986 (Lepidoptera: Tortricidae: Sparganothini) Author Kruse, James J. Author Powell, Jerry A. text Zootaxa 2009 2009-07-06 2150 1 1 78 https://biotaxa.org/Zootaxa/article/view/zootaxa.2150.1.1 journal article 10.11646/zootaxa.2150.1.1 1175­5334 5311432 12. Sparganothoides lugens (Walsingham) Figs. 18 , 45 , 72 Sparganothis lugens Walsingham 1913: 223 . Sparganothoides lugens ; Powell 1986: 376 ; Powell et al. 1995: 149 ; Brown 2005: 564 . Diagnosis . The dark brownish gray forewing, with distinct basal/subbasal, median, and subapical fasciae, distinguishes this species from its congeners. The unusual shape of the valva is also distinct. Description . Male . Head : Frons with white-tipped gray to brownish gray scales produced into a weak hood; vertex roughened, with white-tipped gray to brownish gray scales. Labial palpus pale brownish gray mesally, white-tipped gray to brownish gray scales laterally. Antennal scaling gray to dark gray. Thorax : Dorsum smooth scaled, gray to brownish gray. Forewing length 8.1–9.2 mm (= 8.5; n = 9). Forewing lacking costal fold; interfascial regions pale gray, gray, and brownish orange, distinct basal/subbasal median and subterminal fasciae dark grayish brown, well defined at costa, slightly weakened toward dorsum; dark grayish brown transverse strigulae throughout subterminal and terminal areas. Fringe pale orange to yellowish white. Hindwing gray, with dense gray mottling throughout in some specimens. Abdomen : Genitalia ( Fig. 18 ; slide #JAP3175; EME; Mexico , Veracruz , 1 mi W Fortín; n = 7) with uncus long, slender, uniform in width, elbowed at nearly 90° angle at two-thirds distance from base, with long setae dorsally, long lobes laterally and subapically, small setae on lobes ventrally; tegumen raised, squarish at base of uncus; socius rounded posteriorly, secondary arms long, slender, abruptly angled near middle, enlarged apices asymmetrically bilobed, mitten-shaped, thumb portion narrow, widening distally; transtilla sclerotized, straight, spines short, numerous over most of posterior margin, lacking median lobe; valva elongate, subrectangular, without sclerotized crease, costa curved, sacculus sinuate, pulvinus absent; phallus pistol-shaped, aedeagus tapered and slightly curved, shorter than phallobase, attenuate apically, attached to juxta by strongly sclerotized process; cornuti with minute spine near base. Female . Head, Thorax : Essentially as described for male. Forewing length 8.8–11.1 mm (= 10.1; n = 10). Abdomen : Genitalia ( Fig. 45 ; slide #JAP3152; NMNH; Mexico , Oaxaca , Salina Cruz ; n = 5) with papillae anales parallel-sided, subtriangular posteriorly; sterigma narrow, strongly sclerotized ventrally, concave anteriorly, weakly bilobed; ductus bursae short, widening anteriorly; corpus bursae large, irregularly rounded; signum ca. three times as long as wide, simple, curved, weakly bilobed, blunt at apices. Type material . Holotype : Female: MEXICO : VERACRUZ : Jalapa, [no date], M. Trujillo ( BMNH ). Paratypes (2M). MEXICO : VERACRUZ : Jalapa , [no date] (1 M; lacking abdomen), [no collector] ( BMNH ), [no date] ( 1♂ ), [no collector] ( NMNH ) . Additional specimens examined ( 10♂ , 11♀ ). COSTA RICA : PUNTARENAS : Monteverde , 8–10.vi.1986 ( 2♀ ), J. Chemsak & H. Katsura ( EME ) , 1400 m , 29–31.iii.1992 ( 1♀ ), S. McCarty & J. Powell ( EME ) . MEXICO : OAXACA : Salina Cruz , [no date] ( 2♀ ), W. Schaus ( NMNH ) . VERACRUZ : Jalapa , [no date] ( 1♂ , 1♀ ), M. Trujillo ( BMNH ) , [no date] ( 1♂ ), M. Trujillo ( NMNH ) . Orizaba , [no date] ( 1♂ ), W. Schaus ( NMNH ) . N. Chocoman , 30.vii.1966 ( 1♀ ), O. Flint & Ortiz ( NMNH ) . 1 mi W Fortín , 25.xii.1963 , collected as larva on “misc. leaves”, reared on lettuce, emgd. 20.ii.1964 ( 1♂ ), C. O’Brien ( EME ) . 4 km SW Fortín , 900 m , 12.viii.1990 ( 1♂ ), J. Doyen ( EME ) . Río Metlec , Fortín de las Flores , 15.viii.1987 ( 1♂ ), J. Brown ( EME ) . Near Huatusco , 25–26.vii.1965 (1 F), O. Flint & Ortiz ( NMNH ) . Near San Andres , 5 km W Jalapa , 1600 m , 19.viii.1987 ( 2♀ ), J. Brown & J. Powell ( EME ) . Las Minas , SE Las Vigas , 18.viii.1987 ( 1♀ ), J. Doyen ( EME ) . Coscomatepec , 1.viii.1975 ( 1♂ ), T. Taylor ( LACM ) . 5 km S Coscomatepec , 8.vi.1991 ( 1♂ ), J. Chemsak ( EME ) . 6 km S Coscomatepec , 1370 m , 12.viii.1987 ( 2♂ ), J. Brown & J. Powell ( EME ) . Immature stages . One small larva was found in December by C.W. O’Brien feeding on “miscellaneous leaves” and was reared on lettuce. Biology . Adults have been collected from early June through mid-August in southern Mexico , and in March and June in Costa Rica . Distribution . Sparganothoides lugens has been recorded from southern Mexico to Costa Rica . Remarks . Females from Costa Rica are grayer, the papillae anales are more rounded posteriorly, and male cornuti deposited in the ductus bursae are wider, less curved, and possess a larger spine at the base than those from Mexico . These features suggest that specimens from Costa Rica may not be conspecific with those from Mexico .