Revision of the carnivorous snail genus Discartemon Pfeiffer, 1856, with description of twelve new species (Pulmonata, Streptaxidae) Author Siriboon, Thanit Author Sutcharit, Chirasak Author Naggs, Fred Author Rowson, Ben Author Panha, Somsak text ZooKeys 2014 401 45 107 http://dx.doi.org/10.3897/zookeys.401.7075 journal article http://dx.doi.org/10.3897/zookeys.401.7075 1313-2970-401-45 03F455BB9AA64D5DA89246C6A6D3F42A 03F455BB9AA64D5DA89246C6A6D3F42A 27. Discartemon conicus Siriboon & Panha sp. n. Figs 10E, F, 23, Table 3 Type material. Holotype CUMZ 6255 (Fig. 10E). Measurement: shell height 4.5 mm, shell width 7.2 mm, and with 6 whorls. Paratypes: CUMZ 6033 (2 shells) from the type locality. Type locality. Gau Cerita, Langawi, Malaysia, 6°27'21.8"N , 99°49'29.8"E . Diagnosis. This species differs from Discartemon roebeleni , Discartemon sangkarensis , Discartemon vandermeermohri and Discartemon kotanensis sp. n., in having a smaller shell, higher spire, a nearly smooth shell surface, an angular last whorl, a sub-quadrangular aperture with a sinulus, and in having only one parietal lamella. Description. Shell. Shell globose-heliciform, white and translucent; whorls 6, spire elevated conical, with distinct suture. Shell surface glossy, smooth with thin transverse ridges near aperture; varices present. Embryonic shell large, about 21/2 whorls, with a smooth surface; following whorls regularly coiled. Last whorl angular, inflated and regularly expanded. Umbilicus open and deep. Aperture sub-quadrangular with sinulus; peristome discontinuous, expanded and reflected. Apertural dentition of only one parietal lamella (Fig. 10E). Distribution. Known only from the type locality among limestone karsts up to 100 meters amsl, surrounded by mangrove forests on the northeast Langkawi Island coastline. Remarks. The new species is apparently rare and extensive searching yielded only three examples. Figure 11. Genitalia and pallial complex. A-C Discartemon discus CUMZ 6257, from Vietnam A reproductive system B insertion of vas deferens into penial sheath, and C circular and excretory systems and mantle edge D, E Discartemon nummus CUMZ 6208, from Patthalung D reproductive system, and E insertion of vas deferens into penial sheath. Figure 12. Genitalia. A, B Discartemon discadentus sp. n. paratype CUMZ 6209 A reproductive system, and B insertion of vas deferens into penial sheath C, D Discartemon hypocrites topotype CUMZ 6199 C reproductive system, and D insertion of vas deferens into penis sheath. Figure 13. Genitalia. A, B Discartemon leptoglyphus CUMZ 6007, from Ipoh, Perak, Malaysia A reproductive system, and B insertion of vas deferens into penial sheath C-F Discartemon afthonodontia sp. n. paratype CUMZ 6210 C reproductive system D insertion of vas deferens into penial sheath E inflation of penis, and F details of hermaphroditic duct and seminal vesicle. Figure 14. Genitalia. A, B Discartemon epipedis sp. n. paratype CUMZ 6215 A reproductive system, and B insertion of vas deferens into penis sheath C, D Discartemon flavacandida sp. n., paratype CUMZ 6216 C reproductive system, and D insertion of vas deferens into penis sheath. Figure 15. Genitalia. A-C Discartemon roebeleni topotype CUMZ 6217 A reproductive system B insertion of vas deferens into penial sheath, and C details of hermaphroditic duct and seminal vesicle D-F Discartemon kotanensis sp. n. paratype CUMZ 6230 D reproductive system, E insertion of vas deferens into penis sheath, and F details of hermaphroditic duct and seminal vesicle. Figure 16. Genitalia. A-C Discartemon megalostraka sp. n., paratype CUMZ 6233 A reproductive system B insertion of vas deferens into penial sheath, and C details of hermaphroditic duct and seminal vesicle D-F Discartemon triancus sp. n., paratype CUMZ 6236 D reproductive system E insertion of vas deferens into penis sheath, and F details of hermaphroditic duct and seminal vesicle. Figure 17. Internal sculpture of genitalia. A-E Discartemon nummus CUMZ 6208, from Patthalung A details of atrium surface B low magnification shows arrangement of penial hooks C high magnification of penial hooks D top view of penial hooks, white arrow indicate tip of hook, and E arrangement of vaginal folds F-I Discartemon discadentus sp. n., paratype CUMZ 6209 F details of atrium surface G high magnification of penial hooks with (inset) shows in low magnification H top view of penial hook, and I arrangement of reticulated vaginal folds. Figure 18. Internal sculpture of genitalia. A-E Discartemon hypocrites , topotype CUMZ 6199 A details of atrial pore on atrium surface B arrangement of penial hooks with high magnification, and (inset) shows in low magnification, C penial hooks with (inset) shows top view of the hook D penial hooks with (inset) shows lateral view of the hook, and E arrangement of vaginal folds F-L Discartemon leptoglyphus CUMZ 6007, from Ipoh, Perak, Malaysia F details of atrium surface G arrangement of penial hooks with (inset) low magnification H high magnification of penial hooks I top view of penial hook J low magnification shows lateral view of penial hooks K high magnification shows lateral view of penial hooks, and L details of vaginal surface. Figure 19. Internal sculpture of genitalia. A-E Discartemon afthonodontia sp. n. paratype CUMZ 6210 A details of atrial pore on the atrium surface B high magnification shows arrangement of penial hooks C top view of penial hook D lateral view of penial hook, and E details of vaginal surface F-I Discartemon epipedis sp. n. paratype CUMZ 6215 F details of the atrium surface G scattered arrangement of penial hooks with high magnification, and (inset) shows in low magnification H top view of penial hook, and I details of vaginal surface J-N Discartemon flavacandida sp. n. paratype CUMZ 6216 J details of atrial pore on the atrium surface K high magnification shows arrangement of penial hooks with top view of penial hook in white square L top view of penial hook (from white square in K) M lateral view of penial hook, and N details of vaginal surface. Figure 20. Internal sculpture of genitalia. A-E Discartemon roebeleni topotype CUMZ 6217 A details of atrium surface B low magnification shows arrangement of penial hooks C lateral view of penial hook D top view of penial hook, and E arrangement of vaginal folds F-J Discartemon kotanensis sp. n. paratype CUMZ 6230 F details of atrial pore on the atrium surface G low magnification shows arrangement of penial hooks H high magnification of penial hooks with top view of penial hook in white square I top view of penial hook (from white square in H), and J arrangement of vaginal folds. Figure 21. Internal sculpture of genitalia. A-F Discartemon megalostraka sp. n. paratype CUMZ 6233 A details of atrial pore on the atrium surface B high magnification of penial hooks with top view of penial hook in white square C top view of penial hook (from white square in B) D low magnification shows arrangement of penial hooks E lateral view of penial hook, and F arrangement of vaginal folds G-L Discartemon triancus sp. n. paratype CUMZ 6236 G details of atrial pore on the atrium surface with (inset) high magnification of atrial pore H high magnification of penial hooks with top view of penial hook in white square, I top view of penial hook (from white square in H) J arrangement of penial hooks K lateral view of penial hooks, and L arrangement of vaginal folds. Figure 22. Radula morphology of A Discartemon discus CUMZ 6257, from Da Nang, Vietnam B Discartemon nummus CUMZ 6208, from Patthalung C Discartemon hypocrites , topotype CUMZ 6199 D Discartemon afthonodontia sp. n., paratype CUMZ 6210 E Discartemon roebeleni , topotype CUMZ 6217 F Discartemon kotanensis sp. n. paratype CUMZ 6230 G Discartemon megalostraka sp. n. paratype CUMZ 6233 H Discartemon triancus sp. n. paratype CUMZ 6236. Figure 23. Approximate locations of the type locality of Discartemon species. (1) Discartemon discus , (2) Discartemon planus , (3) Discartemon sykesi , (4) Discartemon nummus , (5) Discartemon khaosokensis , (6) Discartemon discadentus sp. n., (7) Discartemon discamaximus sp. n., (8) Discartemon circulus sp. n., (9) Discartemon deprima sp. n., (10) Discartemon expandus sp. n., (11) Discartemon plussensis , (12) Discartemon hypocrites , (13) Discartemon leptoglyphus , (14) Discartemon platymorphus , (15) Discartemon afthonodontia sp. n., (16) Discartemon epipedis sp. n., (17) Discartemon flavacandida sp. n., (18) Discartemon lemyrei , (19) Discartemon roebeleni , (20) Discartemon collingei , (21) Discartemon stenostomus , (22) Discartemon sangkarensis , (23) Discartemon vandermeermohri , (24) Discartemon kotanensis sp. n., (25) Discartemon megalostraka sp. n., (26) Discartemon triancus sp. n., and (27) Discartemon conicus sp. n.