Review of Europiella Reuter (Hemiptera: Heteroptera: Miridae: Phylinae: Phylini) from Korea, with a description of a new genus
Author
Duwal, Ram Keshari
Author
Jung, Sunghoon
Author
Lee, Seunghwan
text
Zootaxa
2014
3795
3
383
393
journal article
45796
10.11646/zootaxa.3795.3.9
60c4b804-d3a1-420d-bbbb-7505706a3e99
1175-5326
225539
5B32C6B9-1466-4367-98AD-2E40F40B9351
Europiella artemisiae
(
Becker, 1864
)
Figures: 1A, 2A–B, 3A, 4A–E
Capsus artemisiae
Becker, 1864
: 487
(n. sp.)
Plagiognathus solani
Matsumura, 1917
: 432
(n. sp.) [syn. by Schuh et al., 1995]
Plagiognathus albipennis antennaria
Stichel, 1934
: 282
(n. form)
Europiella artemisiae
: Schuh et al., 1995: 385
(n. syn., figs.); Schuh, 1995: 313 (cat.); 2001: 251 (n. syn.);
Yasunaga, 2001
: 162 (diag., fig.);
Kerzhner and Josifov, 1999
: 344 (cat.).
Diagnosis
. Generally distinguished by small, oval body; variously colored (pale brown, brown to dark black,); entirely dark dorsum with pale areas on either inner lateral side of eyes, mesoscutum and base of cuneus; length of labium nearly equal to length of femora; width of vertex equal to length of antennal segment IV; pale (or dirty yellow) fore and middle legs, and fuscous (or black) metafemora (
Fig. 3
A).
Male genitalia
(
Fig. 4
A–D).
Endosoma
: Shape S–like, slightly twisted mesially, and provided with two apical processes, short process somewhat curved backward as in figure 4A.
Phallotheca
: As in figure 4B.
Left paramere
: As in figure 4D.
Right paramere
: As in figure 4C.
Female genitalia
(
Fig. 4
E). Bursa copulatrix somewhat rectangular, central larger area occupied with semi–sclerotized plates furnished with minute spines; sclerotized rings asymmetrical, anteriorly oval and posterior margin more or less straight.
Measurements.
(
5♂
/ 5♀). Body length 2.69–3.19/ 2.68–3.08; head width across eyes 0.65–0.70/ 0.62–0.70; vertex width 0.29–0.35/ 0.34–0.40; lengths of antennal segments I–IV: 0.23–0.26, 0.77–0.95, 0.59–0.66, 0.37–.041/ 0.21–0.25, 0.73–0.80, 0.48–0.58, 0.33–0.42; total length of labium 1.01–1.19/ 1.10–1.17; mesal pronotal length 0.41–0.45/ 0.43–0.48; basal pronotal width 0.88–1.01/ 0.92–1.03; width across hemelytron 1.10–1.24/ 1.16–1.29; and lengths of metafemur, tibia and tarsus: 0.88–1.09, 1.44–1.63, 0.43–0.53/ 0.93–1.09, 1.42–1.57, 0.39–0.46.
Specimens examined
.
North Korea
:
3♂
, 1♀, Dzikha–ri,
16 km
, Südl. Chongdzin,
31.viii.1970
, M. Josifov.
South Korea
:
Chungcheongbuk–do
:
4♂
, 3♀, Okcheon–gun, Annemyeon, Hyeon–ri,
30.vii.2005
, on light trap, S.H. Lee.
Chungcheongnam–do
:
1♂
, 1♀, Boryeong–si, Ocheon–myeon, Wonsando–ri,
5.viii.2009
, R.K. Duwal;
3♂
, 4♀, Danjin–gun,
9.x.2008
, R.K. Duwal and S. Jung;
3♂
, 1♀, Danjin–gun, Seongmun–myeon, Nanjido–ri,
18.viii.2009
, R.K. Duwal and S. Jung;
1♂
, Danjin–gun, Myeoncheon–myeon, Seongsang–ri,
12.x.2006
, S.H. Lee;
2♂
, 3♀, Seosan–si, Unsan–myeon, Anho–ri,
31.viii.2006
, S.H. Lee;
2♂
, Taean–gun, Iwon–myeon, Nae–ri,
31.viii.2006
;
10♂
, 5♀, Yesan–gun, Sinam–myeon, Yonggung–ri,
3.vii.2006
, J.W. Seong;
1♂
, 1♀, Yesan–gun, Deoksan–myeon, Sacheon–ri, Sudeoksa,
11.vii.1991
, collector unknown.
Gangwon–do
: 1♀, Gangneung–si, Myeongju–dong, Songnim–ri, Yeongokchaen,
27.v.1993
, D.S. Gu;
4♂
, 4♀, Honcheon–gun, Sambong Natural forest,
27.vi.2003
, J.W. Seong; 2♀, Hongcheon–gun, Nae–myeon, Changchon–ri,
5.vii.2007
, Y.J. Lee;
4♂
, 4♀, Wonju–si, Munmak–eup,
1.V.2009
, R.K. Duwal and S. Jung;
6♂
, 2♀, same data as above,
27.V.2009
;
4♂
, 6♀, Pyeongchang–gun, Daegwanryeiong–myeon, Hoenggye 3–ri,
11.viii.2006
, J.W. Seong;
1♂
, Pyeongchang–gun, Jinbu–myeon, Dongsan–ri,
30.vii.2007
, J.W. Seong; 1♀, Taebaek–si, Mt. Hambaek,
14.ix.1999
, G.S. Lee (NAAS);
1♂
, Yeongwol–gun, Sangdong–eup, Naedoek–ri,
24.V.2001
, J.Y. Choi.
Gyeongsangbuk–do
: 1♀, Andong–si, Seokdong–dong,
6.VI.2008
, J.O. Lim; 1♀, Andong–si, Yean–myeon, Ingye–ri,
6.vi.2008
, J.O. Lim;
1♂
, Mungyeong–si, Buljeong–dong, Buljeonsa,
9.viii.2007
, J.W. Seong;
1♂
, Yeongyang–gun, Cheonggi–myeon, Jeongjok–ri,
7.vi.2008
, J.O. Lim.
Gyeonggi–do
: Annyang–si, Gwanak Arboretum,
9.vii.2008
, on light trap, R.K. Duwal and S. Jung;
1♂
, 3♀, Icheon–si, Mt. Manhyeong,
1.vii.2008
, R.K. Duwal and S. Jung;
2♂
, Yangpyeong–gun, Mt. Yongmun,
11.vi.2009
, on light trap, R.K. Duwal and S. Jung;
2♂
, same data as above, on light trap,
18.viii.2009
;
1♂
, 3♀, Incheon, Gangwa–gun, Hajeom–myeon, Mangwol–ri,
1.vii.2008
, R.K. Duwal and S. Jung;
1♂
, 1♀, Gapyeong–gun, Mt. Yumyeong,
14.vi.1997
, S.B. Ahn;
1♂
, Goyang–si, Deokyanggu, Gwansandong,
19.vii.2008
, S.W. Park;
2♂
, 1♀, Incheon, Ganghwa–gun, Yangdo–myeon, Samheung–ri,
10.vi.2007
, J.W. Seong; 1♀, Gwanju–si, Docheok–myeon, Sangnim–ri, Mt. Taehwa,
25.ix.2003
, Jung and Chansik;
1♂
, Suwon, NAAS,
25.vii.1997
, J.Y. Choi (NAAS);
2♂
, Yangpyeong–gun, Gangsangmyeon, Sewol–ri,
30.vi.2008
, R.K. Duwal and S. Jung.
Gyeongsangnam–do
:
1♂
, Geoje–si, Geoje Arboretum,
25–27.viii.2008
, on
Artemisia
sp., R.K. Duwal and S. Jung;
7♂
, 1♀, Goeje–si, Irun–myeon, Jisimdo,
28–29.viii.2008
, on
Artemisia
, R.K. Duwal and S. Jung.
Jeju–do
:
3♂
, Jeju–si, Mt. Heandong, Eorimog
27.viii.1997
, S.B. Ahn;
1♂
, Jeju–si, Gujwa–eup, Gimnyeong–ri,
27.viii.1997
, S.B. Ahn; Jeju–si, Mt. Haeandong, Eorimog,
27.VIII.1994
, D.S. Gu; 2♀, Jeju–si, Aeweol–eup, Eoeum–ri,
14.v.2003
;
24♂
, 12♀, Jeju–si,
12–15.v.2008
, on
Artemisia
sp., T. Yasunaga, R.K. Duwal and S. Jung; 1♀, Seogwipo–si, Andeok,
15.v.2003
, J.W. Seong.
Jeollanam–do
:
2♂
, 1♀, Gwangyang–si, Ongnyong–myeon, Chusan–ri,
16–19.vi.2008
, R.K. Duwal and S. Jung; 1♀, Hwasun–gun, Hwasun–eup, Suman–ri,
12.ix.1996
, M.L. Lee.
Japan
:
Hokkaido:
2♂
, 2♀, Yunotai, Hiyama,
19.viii.1994
, T. Yasunaga, det. by T. Yasunaga, 2008;
1♂
, Higashikawa town,
22.viii.1998
, on
Artemisia
spp. T. Yasunaga; 5th instar, Tobetsu town,
28.vi.1998
, T. Yasunaga.
Distribution
. Asia: [
Azerbaijan
,
China
,
Japan
,
Kazakhstan
,
Korea
*,
Russia
,
Uzbekistan
], North
America
, and European continents.
Host
.
Artemisia absinthium
L. and
A
.
vulgaris
L. (
Asteraceae
) [Wagner, 1975].
Remarks
.
Europiella artemisiae
is the most common species of the genus occurring in
Korea
, feeding on
Artemisia
, and abundantly emerging from early spring to late autumn. The formerly reported species from
Korea
(
Kwon et al., 2001
),
Europiella albipennis
is considered a misidentification of
E
.
artemisiae
, because all the collected materials during our survey from
2008–2011
, and the North Korean materials (collections of Josifov) which were observed in this study, were identified as
E
.
artemisiae
rather than
E
.
albipennis
. Therefore, we confirm
E
.
artemisiae
occurs in
Korea
; and
E
.
albipennis
is removed from the Korean list.