Evolutionary history of the devilrays (Chondrichthyes: Myliobatiformes) from fossil and morphological inference Author Adnet, Sylvain Author Cappetta, Henri Author Guinot, Guillaume Author Sciara, Giuseppe Notarbartolo Di text Zoological Journal of the Linnean Society 2012 2015-07-06 166 1 132 159 http://dx.doi.org/10.1111/j.1096-3642.2012.00844.x journal article 10.1111/j.1096-3642.2012.00844.x 0024-4082 5408589 ARGOUBIA GEN. NOV. Type horizon: Samlat Formation, Gerran member ( Ratschiller, 1967 ) – late Middle Eocene/Late Eocene. Etymology: From the close souk of El Argoub. Type species: Argoubia barbei gen. et sp. nov. Occurrences: Only known from the type locality, Late Eocene. Diagnosis: Mobulid only known by isolated teeth of small size. The crown is much higher than the root. Occlusal face rather flat and less developed transversally than the base of the crown in occlusal view. Some smooth and little salient folds labiolingually disposed. The lingual margin of this face is irregularly cut out. The labial face of the crown is oblique in profile and is strongly ornamented, with vertical folds bearing many small spines or tubercles irregularly disposed. These folds are separated by deep grooves. The ornamentation of the lingual face is very similar to that of the labial face, but with fewer spines and tubercles. The basal bulge is thick but not very salient. The lower part of the labial visor is oblique and well developed. The root is not high, with three to four lobes separated by broad and deep furrows. Their basal face is flat with marginal lobes of triangular outline.