A new species of Quercus genus (Fagaceae) from Son Tra Peninsula, Central Vietnam
Author
Ngoc, Nguyen Van
https://orcid.org/0000-0002-1518-9470
Faculty of Biology, Dalat University, 01 - Phu Dong Thien Vuong, Dalat, Vietnam
mr.ngocnv@gmail.com
Author
Binh, Hoang Thi
https://orcid.org/0000-0002-0829-2955
Faculty of Biology, Dalat University, 01 - Phu Dong Thien Vuong, Dalat, Vietnam
Author
Son, Hoang Thanh
https://orcid.org/0000-0003-0351-929X
Department of Forest Phytodiversity, Silviculture Research Institute, Vietnamese Academy of Forest Sciences, Hanoi, Vietnam
Author
Suyama, Yoshihisa
https://orcid.org/0000-0002-3136-5489
Field Science Center, Graduate School of Agricultural Science, Tohoku University, 232 - 3 Yomogida, Naruko-onsen, Osaki, Miyagi 989 - 6711, Japan
Author
Yahara, Tetsukazu
https://orcid.org/0000-0001-5105-7152
Kyushu Open University, 744 Motooka, Fukuoka, 819 - 0395, Japan
text
PhytoKeys
2022
2022-08-26
206
61
73
http://dx.doi.org/10.3897/phytokeys.206.85635
journal article
http://dx.doi.org/10.3897/phytokeys.206.85635
1314-2003-206-61
A0DEE4918D5B5E54BF693D11AA8BE230
Quercus sontraensis Ngoc, H.T.Binh & Son
sp. nov.
Fig. 4
Diagnosis.
Quercus sontraensis
is morphologically similar to
Q. cambodiensis
and
Q. langbianensis
in leaf shape (lanceolate to elliptic), cupules with 6-9 rings and cupule enclosing less than
1/2
of the nut. However,
Q. sontraensis
is distinguished from
Q. cambodiensis
by its leaf margin regularly and distinctly serrate on the upper 1/3-1/4 (-1/5) of the lamina (vs. almost entire or with a few low teeth in upper 1/4), bowl-shaped cupule (vs. cup-shaped), cupule bract margin distinctly toothed in all rings, except two upper rings (vs. entire, except distinctly toothed in two lower rings) and differs from
Q. langbianensis
in having bowl-shaped cupule (vs. cup-shaped), cupule bract margin distinctly toothed in all rings, except two upper rings (vs. distinctly toothed in all rings), broadly ellipsoid nut (vs. obovoid to ellipsoid) and slightly convex nut scar (vs. convex).
Figure 4.
Quercus sontraensis
Binh, Ngoc & Son
A
leafy twig with male inflorescences
B, C
young leaves covered with densely golden hairs
D
mature fruits
E
stylopodium
F, G
side and base of the cupule
H
inside of the cupule
I
nut (lateral view)
K
nut (top view). Materials from
Son et al. QC201
.
Type
.
Vietnam
.
Da Nang
City
,
Son Tra Peninsula
,
Son Tra Nature Reserve
, in evergreen forest, alt.
340 m
,
16°08'22.90"N
,
108°15'28.85"E
,
9 October 2016
,
Son H.T.
,
Binh H.T.
,
Ngoc N.V.
QC201
(
holotype
DLU!, isotypes HN!, VNM!, VAFS!)
.
Description.
Tree, 12 m tall. Terminal and lateral buds ovoid, 3-5 mm long, 1.5-3 mm in diam., scales in 3-5 rows, imbricate, ovate-triangular, ca. 1.5
x
2 mm, apex obtuse, margin yellowish-brown ciliate, appressed hairy on both surfaces. Young twigs greyish-brown, 1.5-2 mm in diam., densely curly golden hairy, sometimes sulcate, old twigs greyish-brown, glabrous, lenticellate. Leaves alternate; blades elliptic to elliptic-lanceolate, 7-12.5
x
2.2-3.5 cm, acuminate or sometimes acute at apex, cuneate at base, margin regularly and distinctly serrate in the upper 1/3-1/4 (-1/5), densely golden hair on both surfaces when young, glabrescent, midrib sunken adaxially, prominent abaxially, lateral veins (8-)11-14 pairs, straight and running into the teeth of margin, slightly sunken adaxially, prominent abaxially, at an angle of 45-50(-52) degrees from midrib, tertiary veins scalariform, faint on the upper surfaces and conspicuous on the lower surfaces; petioles (0.7-)1-1.5 cm long, densely curly golden hairy when young, soon glabrous. Male inflorescence 7-9 cm long. Female inflorescence 2.5-3.5 cm long, female flowers solitary. Infructescences axillary or terminal, erect, rachis 0.5-0.8 (-1.2) cm long, 3-4 (-6) mm in diam., densely golden hairy. Fruits 1-3, 3-3.3 cm long (including cupule), sessile; cupules bowl-shaped, 1.2-1.7 (-1.9) cm long, 1.3-2.2 cm in diam., enclosing 1/3 of the nut when mature, both outside and inside covered with densely appressed yellowish-brown hairs, wall ca. 1.5-3 mm thick, comprising scales, scales arranged in 7-8 rings, margin of the ring distinctly toothed in all rings, except two upper rings; nut broadly ellipsoid 2.3-2.6 cm long, 1.7-2.0 cm in diam., densely golden hairy, apex nearly flat, densely appressed curly golden hairs around stylopodia, stylopodia up to 1.5 mm long, basal scar 1.1-1.3 cm in diam., slightly convex, glabrous.
Distribution.
Vietnam. Da Nang City, Son Tra Nature Reserve.
Etymology.
The specific epithet is derived from the district name of the type locality, Son Tra Nature Reserve, Son Tra Peninsula, Da Nang City, Central Vietnam.
Vernacular name.
Sồi
Sơn
Tra
(suggested here).
Phenology.
Flowering from January to March, fruiting specimens were collected from September to October.
Additional specimens examined.
Vietnam
.
Da Nang
City
,
Son Tra Peninsula
,
Son Tra Nature Reserve
, in evergreen forest,
385 m
elev.,
16°07'41.7"N
,
108°15'55.7"E
,
20 September 2019
,
Son H.T.
QC202
[fr.] (DLU!, VAFS!); ibid.,
428 m
elev.,
16°07'00.3"N
,
108°17'40.5"E
,
20 September 2019
,
Son H.T.
QC203
[fl.] (DLU!, VAFS!)
.
Preliminary conservation status.
During our floristic survey inside the protected area of Son Tra Nature Reserve, less than 10 mature individuals of
Quercus sontraensis
were found in the evergreen forest, from 340 to 430 m altitude. According to the criteria established by the IUCN Red List (
IUCN 2019
),
Q. sontraensis
is qualified as Critically Endangered (CR), based on the extent of occurrence (EOO 1.47 km2) and area of occupancy (AOO 12.0 km2) [CR B1ab(i,ii,iii) 1 B2ab(i,ii,iii)].