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<mods:title id="C9739349F99EEC67DF5A528CDAE09CB1">A taxonomic synopsis of Heliotropiaceae and new combinations in Heliotropium from Thailand</mods:title>
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<mods:namePart id="1A7726D02C691AAEE5C21B4E348F3C32">Rueangsawang, Kanokorn</mods:namePart>
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<mods:namePart id="729AD24A2F41EDD333282969863CFAA6">Chantaranothai, Pranom</mods:namePart>
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<mods:affiliation id="8B0CFBA1F604188B0E8BF6F3EF30128C">Applied Taxonomic Research Center, Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand &amp; Royal Botanic Gardens, Kew, Richmond, Surrey TW 9 3 AE, UK</mods:affiliation>
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<taxonomicName id="875D023BE1262813A0F25C3C300740BA" ID-CoL="4GDN" ID-ENA="454092" LSID="96E76C6D-2C87-5FB0-BB39-3EB5C95A34CB" authority="Nutt., Trans. Amer. Philos. Soc., ser. 2, 5: 189. 1836." authorityName="Nutt., Trans. Amer. Philos. Soc., ser. 2, 5: 189. 1836." class="Magnoliopsida" family="Heliotropiaceae" genus="Euploca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Euploca" order="Boraginales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="genus">Euploca Nutt., Trans. Amer. Philos. Soc., ser. 2, 5: 189. 1836.</taxonomicName>
</paragraph>
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<paragraph id="4EE23E11D1EEDE87A899FD47F8846F19" pageId="0" pageNumber="189">
<taxonomicName id="FA019B875A6B2F6F8635F53C9819971D" authorityName="Rueangsawang &amp; Chantaranothai" authorityYear="2023" class="Magnoliopsida" family="Heliotropiaceae" genus="Heliotropium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Heliotropium" order="Boraginales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="genus">Heliotropium</taxonomicName>
[unranked] 
<taxonomicName id="B9021D8607AC51687F3D3CD6079648EA" class="Magnoliopsida" family="Heliotropiaceae" genus="Orthostachys" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Orthostachys" order="Boraginales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="genus">Orthostachys</taxonomicName>
R.Br., Prodr. Fl. Nov. Holland.: 493. 1810. Type species: 
<taxonomicName id="6E32BF0CD4221DB810745FCE0787FEC6" class="Magnoliopsida" family="Heliotropiaceae" genus="Heliotropium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Heliotropium foliatum" order="Boraginales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="species" species="foliatum">Heliotropium foliatum</taxonomicName>
R.Br.
</paragraph>
<paragraph id="552BBF5FEC4BB6A5125F72E8AF42240B" pageId="0" pageNumber="189">
<taxonomicName id="B1E783B5904F9537560FE07ECB547C48" authorityName="C.F.P.Martius" authorityYear="1827" class="Magnoliopsida" family="Heliotropiaceae" genus="Preslaea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Preslaea" order="Boraginales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="genus">Preslaea</taxonomicName>
Mart., Nov. Gen. Sp. Pl. 2: 75. 1827, nom. illeg.
</paragraph>
<paragraph id="4861AE4FA19F037199B543564F046C84" pageId="0" pageNumber="189">
<taxonomicName id="6BEE501B53A547BB24B1F7341BAE165C" class="Magnoliopsida" family="Heliotropiaceae" genus="Heliotropium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Heliotropium subsp. sect." order="Boraginales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="section" section="Orthostachys">Heliotropium sect. Orthostachys</taxonomicName>
(R.Br.) G.Don, Gen. Syst. 4: 361. 1838. Type species: Based on 
<taxonomicName id="3CDD564E3A986D89A2FF89AADBAA02B1" authorityName="Rueangsawang &amp; Chantaranothai" authorityYear="2023" class="Magnoliopsida" family="Heliotropiaceae" genus="Heliotropium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Heliotropium" order="Boraginales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="genus">Heliotropium</taxonomicName>
[unranked] 
<taxonomicName id="2D124C6376648F985056142A57EA77BE" class="Magnoliopsida" family="Heliotropiaceae" genus="Orthostachys" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Orthostachys" order="Boraginales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="genus">Orthostachys</taxonomicName>
R.Br.
</paragraph>
<paragraph id="D376F0B919C2A921ECB89AB1E47830CE" pageId="0" pageNumber="189">
<taxonomicName id="ABC6923143A1EF1BC2BCD827235280E3" class="Magnoliopsida" family="Heliotropiaceae" genus="Schleidenia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Schleidenia" order="Boraginales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="genus">Schleidenia</taxonomicName>
Endl., Gen. Pl.: 646. 1839. Type species: 
<taxonomicName id="081854B499E78DC8B54BCB74C6376346" class="Magnoliopsida" family="Heliotropiaceae" genus="Preslaea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Preslaea paradoxa" order="Boraginales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="species" species="paradoxa">Preslaea paradoxa</taxonomicName>
Mart.
</paragraph>
<paragraph id="28ECCA6A6FE9C84106378D8DC6FC4878" pageId="0" pageNumber="189">
<taxonomicName id="35A17723ED546E0D66AC5B3790F8103A" class="Magnoliopsida" family="Heliotropiaceae" genus="Orthostachys" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Orthostachys" order="Boraginales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="genus">Orthostachys</taxonomicName>
(R.Br.) Spach, Hist. Nat. 
<normalizedToken id="4DB659DD56B5971888324D91EBCF5596" originalValue="Vég">Veg</normalizedToken>
. 9: 32. 1840. Type species: Based on 
<taxonomicName id="49821CDDE9564A322BD203F7C76688AF" authorityName="Rueangsawang &amp; Chantaranothai" authorityYear="2023" class="Magnoliopsida" family="Heliotropiaceae" genus="Heliotropium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Heliotropium" order="Boraginales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="genus">Heliotropium</taxonomicName>
[unranked] 
<taxonomicName id="0A352B3384F2549F7643F4BC05A2565E" class="Magnoliopsida" family="Heliotropiaceae" genus="Orthostachys" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Orthostachys" order="Boraginales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="genus">Orthostachys</taxonomicName>
R.Br.
</paragraph>
<paragraph id="842F9662FE31D7CF729C414F9302DFE8" pageId="0" pageNumber="189">
<taxonomicName id="D6D12F8C27E840CA2905D3FB360D005D" class="Magnoliopsida" family="Heliotropiaceae" genus="Hilgeria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hilgeria" order="Boraginales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="genus">Hilgeria</taxonomicName>
<normalizedToken id="61192AEB829CA113B3059F8AD9EF4248" originalValue="Förther">Foerther</normalizedToken>
, Sendtnera 5: 132. 1998. Type species: 
<taxonomicName id="8294934CA1A5A72DFD082BB68AEBBB5B" authorityName="Foerther" authorityYear="1998" baseAuthorityName="Urb. &amp; Ekman" class="Magnoliopsida" family="Heliotropiaceae" genus="Hilgeria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hilgeria hypogaea" order="Boraginales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="species" species="hypogaea">Hilgeria hypogaea</taxonomicName>
(Urb. &amp; Ekman) 
<normalizedToken id="B2CA9E12D20204635E3BE58E459C07A8" originalValue="Förther">Foerther</normalizedToken>
.
</paragraph>
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<paragraph id="FF5C043547912D99B8F5B821ADF723F7" pageId="0" pageNumber="189">Type species.</paragraph>
<paragraph id="4F0DBB1E8213F80FB16FD13EFE3FA597" pageId="0" pageNumber="189">
<taxonomicName id="DB0BED18C86463C9D51E6496D97CA3C7" class="Magnoliopsida" family="Boraginaceae" genus="Euploca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Euploca convolvulacea" order="Boraginales" pageId="0" pageNumber="189" phylum="Tracheophyta" rank="species" species="convolvulacea">
<emphasis id="574C9063393F5FC99433A46B72CD3C2F" italics="true" pageId="0" pageNumber="189">Euploca convolvulacea</emphasis>
</taxonomicName>
Nutt.
</paragraph>
</subSubSection>
<subSubSection id="2AF6C0E47F4239CE8811ABAD6D4C13CF" pageId="0" pageNumber="189" type="description">
<paragraph id="E0C6C27A4063FB6E444CA4F0E264C311" pageId="0" pageNumber="189">Description.</paragraph>
<paragraph id="662A4739E136D38B718FA493707C7AE5" pageId="0" pageNumber="189">Annual herbs. Leaves petiolate or sessile; lamina linear to ovate, apex acute to obtuse, margin entire or revolute, pubescent to strigose or silky silver hairs on both surfaces. Inflorescences terminal or axillary, spike-like, scorpioid or subcapitate cymes, bracteate. Flowers 5-merous. Calyx 5-lobed, lobes linear to lanceolate. Corolla white with a yellow-orange throat inside, tubular or funnel-shaped, lobes orbicular to oblong, throat pubescent inside. Stamens included, sessile or with short filaments; anthers, elliptic-oblong. Ovary divided into 4 lobes. Fruits separating into four 1-seeded nutlets.</paragraph>
</subSubSection>
</treatment>
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