Revision of the Mesoamerican species of Calolydella Townsend (Diptera: Tachinidae) and description of twenty-three new species reared from caterpillars in Area de Conservacion Guanacaste, northwestern Costa Rica Author Fleming, AJ Author Wood, D. Monty Author Smith, M. Alex Author Hallwachs, Winnie Author Janzen, Daniel H text Biodiversity Data Journal 2018 6 11223 11223 http://dx.doi.org/10.3897/BDJ.6.e11223 journal article http://dx.doi.org/10.3897/BDJ.6.e11223 1314-2828--11223 Calolydella alexanderjamesi Fleming & Wood sp. n. Materials Type status: Holotype . Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017782 ; recordedBy: D.H. Janzen, W. Hallwachs & gusaneros ; individualID: DHJPAR0017782; individualCount: 1 ; sex: M ; lifeStage: adult ; preparations: pinned; otherCatalogNumbers: 99-SRNP-16175, BOLD:AAW8659, ASTAR493-07; Taxon: scientificName: Calolydellaalexanderjamesi; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: alexanderjamesi; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica ; countryCode: CR; stateProvince: Guanacaste; county: Sector Santa Rosa; locality: Area de Conservacion Guanacaste ; verbatimLocality: Area Administrativa; verbatimElevation: 295 ; verbatimLatitude: 10.8376; verbatimLongitude: -85.6187; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming ; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the erebid moth, Correbiaundulata ; verbatimEventDate: 16-Nov-1999 ; Record Level: language: en; institutionCode: CNC ; collectionCode: Insects ; basisOfRecord: Pinned Specimen Description Male (Fig. 3a, b, c). Length: 7mm. Head (Fig. 3b): frontal setae extending to base of postpedicel; fronto-orbital plate gold, sparsely setulose throughout; parafacial at least 50% silver pollinose. Thorax (Fig. 3a, c): gold pollinose dorsally with a gold tinge on lateral surfaces; outermost two thoracic vittae twice as wide as innermost two, these becoming fused postsuturally; postpronotum with two setae (inner basal seta absent); 3:3 acrostichal setae; 3:4 dorsocentral setae; 2:2 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite bare; scutellar discal setae situated as wide apart as subapical scutellar setae. Wing vein R4+5 with at most three small setulae dorsally, near base. Abdomen (Fig. 3a): ground color dark brown-orange with uninterrupted transverse marginal pollinose bands; abdominal pollinosity gold dorsally, silver ventrally, with an orange spot lateroventrally at base of ST1+2; T3 with one pair of median marginal setae and one pair of discal setae; T4 with one pair of discal setae. Terminalia: not examined. Female: not known at this time. Diagnosis Calolydella alexanderjamesi can be distinguished from all other species of Calolydella by the following combination of traits: parafacial mostly gold, fronto-orbital plate with small black setulae interspersed among frontal setae, anatergite bare, and thoracic pollinosity gold on both dorsal and lateral surfaces. Etymology The specific epithet is in honor of Alexander James of Levittown, Pennsylvania, in recognition of the moral and family support of his mother, Tanya Dapkey, in her efforts curating and preparing ACG parasitoid flies for DNA barcoding. Distribution Costa Rica, ACG, Guanacaste Province, Area Administrativa, 295m. Ecology Calolydella alexanderjamesi has been reared once from Correbia undulata (Druce, 1884) ( Lepidoptera : Erebidae ), in dry forest.