High mitochondrial DNA sequence divergence in New Guinea Carabdytes stream beetles and the taxonomist's dilemma when other evidence is kind of subtle ... (and collecting localities are far far away) Author Skale, Andre Bluecherstrasse 46, 95030 Hof / Saale, Germany Author Taenzler, Rene Department of Entomology, Zoological State Collection, Munich, 81247 Germany Author Hendrich, Lars Department of Entomology, Zoological State Collection, Munich, 81247 Germany Author Balke, Michael Department of Entomology, Zoological State Collection, Munich, 81247 Germany & GeoBioCenter, Ludwig-Maximilians-University, Munich, Germany coleoptera-zsm@zsm.mwn.de text ZooKeys 2012 2012-11-30 247 31 43 http://dx.doi.org/10.3897/zookeys.247.3812 journal article http://dx.doi.org/10.3897/zookeys.247.3812 1313-2970-247-31 FFDC1A20FFAEFFC86C13FFB6FF9CFFAE 577872 Carabdytes upin tindige ssp.n. Type locality. West Papua, Arfak Mountains, Siyoubrig, 01°06.26'S, 133°54.41'E. Description. Holotype: ♂ (MZB): Indonesia, West Papua, Arfak Mountains, stream near Siyoubrig 01°06.26'S, 133°54.41'E, 1400-1800 m, 24. -28.II.2007, leg. A. Weigel. Paratypes: 2♂♂ 1 ♀ (CSH, ZSM): same locality data as holotype (the female was sequenced). Habitus as in Fig. 3 ; total length: 11.6-12.0 mm; total width: 5.3-5.5 mm. Dark brown to almost black; labrum, lateral margin of pronotum and all body appendages paler reddish brown; elongate. Colour. Head black, labrum reddish brown, clypeus with indistinctly reddish colour almost reaching eyes; head with indistinctly reddish patch on frons. Pronotum black, with indistinctly median reddish patch, posterior angles reddish. Ventral surface blackish. Venter dark brown. Structures. Head with fine, sparse punctation interspersed with coarser punctures between eyes and behind anterior clypeal margin. Pronotum shining, posterior angles with irregular, coarse punctures, lateral margin very strongly. Elytra shining with very indistinctly punctures; each elytra with four rows of coarser, moderately arranged punctures. Lateral wing of metaventrite broad and tongue-shaped; outer margin slightly sinuate; last abdominal ventrite medially emarginate. Legs long and slender. Male. Pro- and mesotarsal claws of similar structure; anterior and posterior claws moderately long and evenly curved; Median lobe of aedeagus relatively slender ( Fig. 3 , shape in Carabdytes upin upin is identical); paramere slender, with distinctly longitudinal striation; setation more or less long ( Fig. 3 ), the setation might be basally shorter in some specimens of Carabdytes upin upin ( Fig. 3 ), but this difference does not appear constant. Diagnosis. Distinguished from Carabdytes upin upin through the molecular and morphological characters mentioned in the results section under "morphology" . With Carabdytes upin upin there is no overall size difference (10.1-12.2 mm). Habitat. Two individuals collected with an aquatic net from the rough gravel at the edge of a stream bed, the stream was rather dry at the time of collection ( Fig. 4 ). The species co-occured with Platynectes spp., Exocelina (= Papuadytes ) spp. and Hydraena cristatigena Jaech & Diaz. Two exemplars were collected with the help of a light trap, approx. 50 m away from the stream. Figure 4. Type locality of Carabdytes upin tindige ssp.n. Distribution. So far known only from the type locality ( Figs 1 , 4 ). Etymology. In loving memory of Samkris "Kris" Tindige, relentless conservationist in Papua, who left us too early. The beetles were collected in the stream bed very close to a birdwatching guesthouse set up by Kris and Shita Prativi above Siyoubrig village .