<document id="7B84F9D27053C9C81B15C5EC0A3CA899" ID-CLB-Dataset="32486" ID-DOI="http://dx.doi.org/10.3897/zookeys.686.12206" ID-GBIF-Dataset="82b32106-bef0-417d-81d4-f47f22c33fa9" ID-PMC="PMC5672565" ID-Pensoft-Pub="1313-2970-686-15" ID-PubMed="29200915" ID-ZooBank="976E23A1CFC74CB381705B59452825A6" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-686-15" ModsDocOrigin="ZooKeys 686" ModsDocTitle="Updated catalogue and taxonomic notes on the Old-World scorpion genus Buthus Leach, 1815 (Scorpiones, Buthidae)" checkinTime="1502744466696" checkinUser="pensoft" docAuthor="Sousa, Pedro, Arnedo, Miquel A. &amp; Harris, D. James" docDate="2017" docId="82B953A10D190217D40F2BBBC694A02A" docLanguage="en" docName="ZooKeys 686: 15-84" docOrigin="ZooKeys 686" docSource="http://dx.doi.org/10.3897/zookeys.686.12206" docTitle="Buthus barcaeus Birula 1909" docType="treatment" docVersion="6" lastPageNumber="25" masterDocId="FF8DFFA394365835FF81FFFFFF80FF91" masterDocTitle="Updated catalogue and taxonomic notes on the Old-World scorpion genus Buthus Leach, 1815 (Scorpiones, Buthidae)" masterLastPageNumber="84" masterPageNumber="15" pageNumber="25" updateTime="1701377877384" updateUser="ExternalLinkService">
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<mods:title id="58F9026BE67CAC9F9E4721C7F93344B0">Updated catalogue and taxonomic notes on the Old-World scorpion genus Buthus Leach, 1815 (Scorpiones, Buthidae)</mods:title>
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. 
<taxonomicName id="55C6FF3DEEC5C8BA2A5A8E44E201562A" ID-CoL="NYYR" authority="Birula, 1909" authorityName="Birula" authorityYear="1909" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus barcaeus" order="Scorpiones" pageId="10" pageNumber="25" phylum="Arthropoda" rank="species" species="barcaeus">Buthus barcaeus Birula, 1909</taxonomicName>
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<paragraph id="226B142AC7DF34C17B99F74A41CC79FC" pageId="10" pageNumber="25">
<taxonomicName id="60D56F1D9D6F3038D806E8D2F82F568C" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus occitanus" order="Scorpiones" pageId="10" pageNumber="25" phylum="Arthropoda" rank="subspecies" species="occitanus" subspecies="barcaeus">Buthus occitanus barcaeus</taxonomicName>
: 
<bibRefCitation id="8C2484400A279317BD93E57D2C07364E" author="Birula, AA" journalOrPublisher="Abteilung fuer Systematik, Geographie und Biologie der Tiere" pageId="23" pageNumber="38" pagination="505 - 522" title="Scorpione und Solifugen von Tripolis und Barka. Nach der Sammlung von Dr. Bruno Klaptocz im jahre 19 - 06. Zoologische Jahrbuecher." volume="28" year="1909">Birula 1909</bibRefCitation>
: 508-511. fig. A, C; 
<bibRefCitation id="4A4EF47CA58AC39E1E65E75B77BC59F2" author="Borelli, A" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="24" pageNumber="39" pagination="148 - 159" title="Contributo allo studio della fauna Libica. Materiali raccolti nelle zone di Misurata e Homs (1912 - 13). Scorpioni." volume="6" year="1914 a">Borelli 1914a</bibRefCitation>
: 155-156; 
<bibRefCitation id="25D5DD69DF521F390FBB1400FB0C93DB" author="Borelli, A" journalOrPublisher="Universita di Torino" pageId="24" pageNumber="39" pagination="1 - 16" title="Missione zoologica del Dr. E. Festa in Cirenaica. XVI. Scorpioni e Solifughi. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R." volume="39" year="1924">Borelli 1924</bibRefCitation>
: 5-7; 
<bibRefCitation id="CC7B4CD9FEB9B0073BBD2AA193D2E8A9" author="Borelli, A" journalOrPublisher="Annali del Museo civico di Storia Naturale Giacomo Doria" pageId="24" pageNumber="39" pagination="346 - 355" title="Risultati zoologici della Missione inviata dalla R. Societa Geografica Italiana per l'explorazione dell'oasi di Giarabud (1926 - 1927). Scorpioni e Solifughi." volume="52" year="1928">Borelli 1928</bibRefCitation>
: 351; 
<bibRefCitation id="795A6CB666FC62163AAF5F41C7E886CD" author="Caporiacco, L di" journalOrPublisher="Annali del Museo civico di Storia Naturale Giacomo Doria" pageId="24" pageNumber="39" pagination="395 - 408" title="Spedizione scientifica all'Oasi di Cufra (Marzo-Luglio 1931). Scorpioni e Solifugi." volume="55" year="1932">Caporiacco 1932</bibRefCitation>
: 395; 
<bibRefCitation id="B7E18917AF5BF0D4AF019D26755A1481" pageId="10" pageNumber="25">Borelli 1934</bibRefCitation>
: 169; 
<bibRefCitation id="2937F94CFFC063D0CC4388D037BEB186" author="Caporiacco, L di" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano" pageId="24" pageNumber="39" pagination="340 - 354" title="Risultati scientifici della Missione del Prof. G. Scortecci nel Fezzan e sui Tassili (1936). Scorpioni e Solifughi." volume="76" year="1937">Caporiacco 1937</bibRefCitation>
: 345; 
<bibRefCitation id="AD0AB4FB230773BBCA052A6D13AF25A7" author="Perez, S" journalOrPublisher="Catedra de Artropodos" pageId="33" pageNumber="48" pagination="1 - 45" title="Un inventario preliminar de los escorpiones de la region Paleartica y claves para la identification de los generos de la region Paleartica Occidental." volume="7" year="1974">
<normalizedToken id="FA2C0511F9CD17DAE6350B9923C96239" originalValue="Pérez">Perez</normalizedToken>
1974
</bibRefCitation>
: 23; 
<bibRefCitation id="8EB79E38FFF070AA039169DB396FF500" author="Levy, G" journalOrPublisher="The Israel Academy of Sciences an d Humanit es, Jerusalem" pageId="30" pageNumber="45" title="Fauna Palaestina. Arachnida I: Scorpiones." year="1980">Levy and Amitai 1980</bibRefCitation>
: 16; 
<bibRefCitation id="97DDC9A769AB829EC595C529EFAA530A" author="El-Hennawy, HK" journalOrPublisher="Serket" pageId="25" pageNumber="40" pagination="95 - 153" title="A catalogue of the scorpions described from the Arab countries (1758 - 1990) (Arachnida: Scorpionida)." volume="2" year="1992">El-Hennawy 1992</bibRefCitation>
: 98, 120; 
<bibRefCitation id="29514DA079165D550AA1EA9DF75F2D67" author="Kovarik, F" journalOrPublisher="Madagaskar, Jihlava" pageId="29" pageNumber="44" title="Stiri." year="1998">
<normalizedToken id="9FFE748D5F1A3B2215853919214D464B" originalValue="Kovařík">Kovarik</normalizedToken>
1998
</bibRefCitation>
: 106; 
<bibRefCitation id="BB7A6153DF90C056A79B1F39F9E9C775" pageId="10" pageNumber="25">Fet and Lowe 2000</bibRefCitation>
: 95; 
<bibRefCitation id="670AD4F8E47A5C49A9089AD1D2359FA6" author="Kovarik, F" journalOrPublisher="Serket" pageId="29" pageNumber="44" pagination="1 - 23" title="A checklist of scorpions (Arachnida) in the collection of the Forschungsinstitut und Naturmuseum Senckenberg, Frankfurt am Main, Germany." volume="8" year="2002">
<normalizedToken id="A18F71DF30E8510BF01F33C5FD27DAB6" originalValue="Kovařík">Kovarik</normalizedToken>
2002
</bibRefCitation>
: 5.
</paragraph>
<paragraph id="57222BFAFE59F68DE2404293C2109799" pageId="10" pageNumber="25">
<taxonomicName id="45D5E22F389FE414EF896B24E4A197FA" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus (Buthus) occitanus" order="Scorpiones" pageId="10" pageNumber="25" phylum="Arthropoda" rank="subspecies" species="occitanus" subGenus="Buthus" subspecies="barcaeus">Buthus (Buthus) occitanus barcaeus</taxonomicName>
: 
<bibRefCitation id="7B2092C06A1E50F4BCF2194B57F73D7B" author="Birula, AA" journalOrPublisher="Horae Societatis Entomologicae Rossicae" pageId="23" pageNumber="38" pagination="115 - 192" title="Ueber Scorpio maurus Linne und seine Unterarten." url="https://www.ntnu.no/ub/scorpion-files/Birula_1910.pdf" volume="39" year="1910">Birula 1910</bibRefCitation>
: 156; 
<bibRefCitation id="FC30A8F0E0FF1008F572843125686BB1" author="Birula, AA" journalOrPublisher="Zapiski Kavkazskogo Muzeya, Seriia A [Memoires du Musee du Caucase, Se ́ rie A]" pageId="23" pageNumber="38" pagination="1 - 253" title="[authored as Byalynitskii-Birulya, A. A.] ArachnoideaArthrogastra Caucasica. Pars I. Scorpiones." volume="5" year="1917 a">Birula 1917a</bibRefCitation>
: 223.
</paragraph>
<paragraph id="24ADE8D8C36E2D6F7E31C8798E79EE2E" pageId="10" pageNumber="25">
<taxonomicName id="11DA10B42AE2E3A22AEC6150A4E96F2C" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus barcaeus" order="Scorpiones" pageId="10" pageNumber="25" phylum="Arthropoda" rank="species" species="barcaeus">Buthus barcaeus</taxonomicName>
: 
<bibRefCitation id="2F1E7D7A5D719C944A4AC038F9CCCE44" author="Kovarik, F" journalOrPublisher="Euscorpius" pageId="29" pageNumber="44" pagination="1 - 16" title="Review of Tunisian Species of the Genus Buthus with Descriptions of Two New Species and a Discussion of Ehrenberg's Types (Scorpiones: Buthidae)." url="http://www.science.marshall.edu/fet/euscorpius/p2006_34.pdf" volume="34" year="2006">
<normalizedToken id="32C91AE18CAC14E9DCA4433897292493" originalValue="Kovařík">Kovarik</normalizedToken>
2006
</bibRefCitation>
: 3, fig. 6; 
<bibRefCitation id="947AA28B846324F062B26725382A378B" author="Kaltsas, D" journalOrPublisher="Journal of Heredity" pageId="28" pageNumber="43" url="http://bozidar-curcic.bio.bg.ac.rs/PDF/16" year="2008">Kaltsas et al. 2008</bibRefCitation>
: 215; 
<bibRefCitation id="1749190B7C826C2A5A24E85258C4ED59" author="Lourenco, WR" journalOrPublisher="Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg" pageId="31" pageNumber="46" pagination="11 - 18" title="A new species of Buthus Leach, 1815 from Egypt (Scorpiones, Buthidae)." volume="16" year="2012">
<normalizedToken id="7814F33C1C3CF4D88AC40840100E3D14" originalValue="Lourenço">Lourenco</normalizedToken>
and Cloudsley-Thompson 2012
</bibRefCitation>
: 15; 
<bibRefCitation id="DC3E19746708568E43F5E85717B41A0D" author="Lourenco, WR" journalOrPublisher="Serket" pageId="31" pageNumber="46" pagination="8 - 15" title="Confirmation of a new species of Buthus Leach, 1815 from Alexandria, Egypt (Scorpiones, Buthidae)." volume="13" year="2012">
<normalizedToken id="E6704952846873913769212B0B4946D5" originalValue="Lourenço">Lourenco</normalizedToken>
and Simon 2012
</bibRefCitation>
: 11; Rossi, Tropea and 
<normalizedToken id="1BEEE42BEE3BDF7991C3B3913BEF9CF8" originalValue="Yağmur">Yagmur</normalizedToken>
2013: 3-5, 7.
</paragraph>
</subSubSection>
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<paragraph id="5240D0DFF22038F60C1A28988A036A9C" pageId="10" pageNumber="25">Type material.</paragraph>
<paragraph id="5B46EC2E24D672290E136AD4A2EB4948" pageId="10" pageNumber="25">4 M, 1 F juv., syntypes (ZIN), Barca (approx. 32.48°, 20.83°), 5 km E from Benghazi (Cyrenaica), Libya.</paragraph>
</subSubSection>
<subSubSection id="6D1BF7DA1D05D42101668291B9849BE4" pageId="10" pageNumber="25" type="distribution">
<paragraph id="681AD1482F132D66FD3C799AE17F2123" pageId="10" pageNumber="25">Distribution.</paragraph>
<paragraph id="A3F6EAE2DB3F8348C637AF0D75FC3D31" pageId="10" pageNumber="25">know from several localities along the Mediterranean coast of Libya.</paragraph>
</subSubSection>
<subSubSection id="03C84F19C8723918923C40794F34865D" pageId="10" pageNumber="25" type="remarks">
<paragraph id="595EF4B8B4197CDDE9FB21278E4E5E4F" pageId="10" pageNumber="25">Remarks.</paragraph>
<paragraph id="87269879734E3C8F9C699C85B743FC4D" pageId="10" pageNumber="25">
The specimens present in the MNHN (F n° 4896), captured in Barca and identified by Vachon in 1974, have no intermediary keel on the fourth metasomal segment, which casts doubts about the use of the character in the diagnosis of 
<taxonomicName id="2570629BC99F2399BE1DA12630D1C434" lsidName="B. barcaeus" pageId="10" pageNumber="25" rank="species" species="barcaeus">B. barcaeus</taxonomicName>
.
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