Systematics and biogeography of the New World species of Trichadenotecnum Enderlein (Insecta: Psocodea: ‘ Psocoptera’: Psocidae) Author Yoshizawa, Kazunori Author Aldrete, Alfonso Neri García Author Mockford, Edward L. text Zoological Journal of the Linnean Society 2008 2008-08-31 153 4 651 723 https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00398.x journal article 10.1111/j.1096-3642.2008.00398.x 0024-4082 5446195 TRICHADENOTECNUM MACULATUM SP. NOV. Material examined: Holotype . Male. Mexico : Nuevo León , 5 km W of Ciénega , hacia Laguna de Sánchez , 1200 m , 25.v.1977 , ANGA ( UNAM ) . Paratypes . 1 female , same data as holotype ; 1 male , 1 female , Nuevo León , 10 mi E Iturbide , Hwy. 60, 13.vi.1962 , J. Campbell ( ISU ) . Etymology: The specific epithet is taken from Latin macula (= spotted), indicating the heavily spotted forewing in this species. Description: Male . Head . White in ground colour; vertical and orbital markings pale brown; coronal suture black; frontal suture very narrowly bordered with pale brown band dorsally; frons with two pairs of pale brown spots ventrally; eye black, IO/D = 1.0; ocelli white, ocellar field black; antennal socket bordered with dark brown band; postclypeus with c . eight rows of pale brown spots at middle, fused with each other ventrally forming a large blackish brown marking, ventro-lateral corner white, lateral margin of postclypeus brown, dorsal margin without marking; anteclypeus brown. Antenna pale brown, pedicel and scape darker. Mouth parts dark brown; third segment of maxillary palpus pale brown. Thorax . Prothorax brown. Mesonotum white; anterior surface of scutum brown; anterior margin of lateral lobe of scutum black; posterior region of scutum with pair of brown markings at middle; scutellum white, median part and anterior margin brown; postnotum brown. Metanotum brown, with pair of whitish oval spots on posterior regions of lobes of scutum, lateral corner white. Meso- and metapleurites brown with longitudinal white band medially. Legs. Coxae dark brown; femora brown except middle part of fore and middle femora white; tibiae pale brown, with darker portion near distal ends; tarsi brown. Forewing ( Fig. 17F ). Densely covered with small spots. Spots in cell a 1 distinct, basal one smaller. Opposing spots in cell r apparent for anterior one but posterior one very small and less distinct. Proximal band broad, strongly constricted around fork of M + Cu, with elongate paler pigmented area in cell CuA. Distal band somewhat indistinct in cell r 5 , band in cell r 3 distinct. Spot on roof of cell m 3 large. Submarginal spots diffuse; spot in cell r 5 larger than others and distinct; spots in cell m 1 , m 2 and m 3 small, sometimes indistinct. Marginal clouds clear, darker along veins. Hindwing. Hyaline, extensively covered with pale brown spots, some spots fused with each other to form a large marking in cell CuP; veins brown. Abdomen . White, each segment with blackish brown band. Terminalia . Clunial arm ( Fig. 25A ) long and narrow, slightly arched, forming pointed free process apically, antero-ventrally with fleshy lobe covered with fine setae. Anterior margin of 8th sternum ( Fig. 25C ) concave; fusion with hypandrium very narrow. Epiproct lobe ( Fig. 25B ) expanded dorsally, wider than long, dorsal margin slightly arched, with rounded projection at middle, posterior surface almost flattened. Paraproct ( Fig. 25A ): ventro-lateral band short, triangular; distal process broad. Hypandrium ( Fig. 25C ) with weakly rugose region laterally; left process sinuate, directed postero-externally at apex; right process of almost equal width from base to apex, projected outward, then strongly bent posterointernally near base, with few denticles at apex; median tongue narrowing medially, then broadened apically, apical margin almost straight, dorsal keel triangular. Phallosome ( Fig. 25D ): pseudoparameres widely separated, longer than wide, directed posteriorly, apical margin rounded or somewhat truncated. Phallobase widest at anterior two-fifths, then narrowing to pointed anterior end. Figure 25. Male terminalia structures of Trichadenotecnum maculatum , showing terminalia in lateral view (A), epiproct in posterior view (B), hypandrium in ventral view (C) and phallosome in ventral view (D). Figure 26. Female genitalia of Trichadenotecnum maculatum , showing subgenital plate (A), gonapophyses (B) and internal plate (C) in ventral view. Length . B 2.6–2.9; FW 3.0–3.7; HW 2.2–2.8. Female . Colour and general morphology almost as in male; eye IO/D = 1.9. Genitalia : Egg guide of subgenital plate ( Fig. 26A ) widened near distal end, postero-lateral corners rounded, apical margin only slightly arched; body of subgenital plate with pair of broad extensions of sclerite antero-medially. Dorsal valve of gonapophyses ( Fig. 26B ) with long distal process; posterior lobe of external valve strongly projected posteriorly. Internal plate as in Figure 26C . Length . B 2.8–3.0; FW 3.2–3.8; HW 2.1–2.9. Distribution: Mexico .