Revision of Eccoptolonthus Bernhauer (Coleoptera: Staphylininae: Philonthina) with descriptions of four new species from China Author Fei, Xu-Dong 0000-0003-0386-5918 Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Rd., Chaoyang District, Beijing 100101, P. R. China. & University of the Chinese Academy of Sciences, 19 A Yuquan Rd., Shijingshan District, Beijing 100049, P. R. China. & feixudong @ ioz. ac. cn; https: // orcid. org / 0000 - 0003 - 0386 - 5918 feixudong@ioz.ac.cn Author Zhou, Hong-Zhang Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Rd., Chaoyang District, Beijing 100101, P. R. China. & University of the Chinese Academy of Sciences, 19 A Yuquan Rd., Shijingshan District, Beijing 100049, P. R. China. text Zootaxa 2021 2021-03-26 4949 3 473 498 journal article 7571 10.11646/zootaxa.4949.3.3 39300142-6cea-4253-80bb-00856fdd5f95 1175-5326 4640469 C22ACA1A-4856-4979-BD7A-DFE37949C2D4 4. Eccoptolonthus eccoptomus ( Tottenham, 1962 ) Tottenham, 1962: 231 ( Philonthus , subgenus Eccoptolonthus , type locality: Natal). Herman, 2001a: 2816 ( Philonthus , catalog); Hromádka, 2010: 497 (transferred to Pseudohesperus , characters); Newton, 2015: 14 (transferred to Eccoptolonthus , catalog). Distribution. South Africa .