The spider family Micropholcommatidae (Arachnida: Araneae: Araneoidea): a relimitation and revision at the generic level
Author
Rix, Michael
Western Australian Museum, Welshpool DC, Perth, Australia
Author
Harvey, Mark
Western Australian Museum, Perth, Welshpool, Australia
text
ZooKeys
2010
2010-02-22
36
36
1
321
journal article
10.3897/zookeys.36.306
4db6b327-7482-432e-a5f6-36f91c79fef3
1313–2970
576620
ADCACC88-6C78-4386-8E33-3F98234ECE92
Taphiassa
robertsi
Rix & Harvey
,
sp. n.
urn:lsid:zoobank.org:act:
D3B8D09B-71C2-49EF-97F1-0C145EEAB3CA
Figs 7P
,
8B, 8D, 8F, 8H
, 10E–F, 152A–B, 152F, 154C–D, 155–162, 215, 223F–G
Type
material.
Holotype
male:
Yanchep National Park
,
Mambibby Cave
(YN-12),
Western Australia
,
Australia
, web strand on rock wall near stair,
31°33'S
,
115°41'E
,
4.VII.1998
,
R. Foulds
(
WAM
T94519
).
Paratypes
:
Allotype
female,
Tuart Cave
(YN-474),
Carabooda area
,
A. Lombardo’s
property,
Western Australia
,
Australia
, off soil cone litter, edge of daylight,
20.III.1999
,
R. Foulds
(
WAM
T94520
)
.
Other
material examined
.
AUSTRALIA
:
Western Australia
:
same data as holotype except
18.V.2002
, 1♁ (
WAM
T94521
SEM♁
)
;
same data as holotype ex-
cept web strand, twilight zone,
30.VI.2001
,
1♀
(
WAM
T94522
)
;
Yanchep National Park
,
Chingah Cave
(YN-416), rock wall under daylight hole, twilight zone,
16.XII.2000
,
R. Foulds
,
1♀
(
WAM
T94523
)
;
Yanchep National Park
, unnamed cave (YN-420),
4 m
into cave, twilight of entrance overhang, dragline,
16.IV.1996
, R.
Foulds
,
1♀
(
WAM
T94524
)
;
Yanchep National Park
,
Carpark Cave
(YN- 18), web strand in rockwall, daylight edge,
7.VII.1998
,
R. Foulds
,
1♀
(
WAM
T94525
SEM
♀
)
;
Bibbulmun Track
, E. of
Albany Wind Farm
, sifting elevated leaf litter,
35°04'10"S
,
117°48'26"E
,
18.III.2008
,
M. Rix
,
M. Harvey
, 1♁ (
WAM
T94529
)
;
D’Entrecasteaux National Park
,
Chesapeake Road
, sifting leaf litter,
34°51'30"S
,
116°24'46"E
,
14.III.2007
,
M. Rix
,
M. Harvey
,
M. Moir
,
1♀
(
WAM
T94528
)
;
Gull Rock National Park
, end of
Ledge Point Road
,
35°00'51"S
,
118°00'23"E
,
17.III.2008
,
M. Rix
,
M. Harvey
, 1♁ (
WAM
T94492
)
;
Limeburners Road
, near
Torndirrup National Park
, sifting elevated leaf litter in deep gully,
35°05'27"S
,
117°54'40"E
,
14.III.2008
,
M. Rix
,
M. Harvey
, 3♁,
1♀
(
WAM
T94527
)
;
Margaret River region
,
Foxhole Cave
(MR-9), from web among rocks in breezeway,
33°55'S
,
115°00'E
,
2.XI.1996
,
R. Foulds
, 1♁ (
WAM
T94526
)
;
Porongurup National Park
,
Waddys Hut
, sifting elevated leaf litter,
34°40'55"S
,
117°50'55"E
,
29.IV.2008
,
M. Rix
,
M. Harvey
, 1♁ (
WAM
T94482
)
;
Sand Patch Beach Reserve
, end of
Mutton Bird Road
, sifting elevated leaf litter,
35°02'41"S
,
117°41'31"E
,
18.III.2008
,
M. Rix
,
M. Harvey
,
1♀
(
WAM
T94530
)
;
Torndirrup National Park
,
Salmon Hole Road
, sifting elevated leaf litter,
35°06'07"S
,
117°58'03"E
,
14.III.2008
,
M. Rix
,
M. Harvey
,
1♀
(
WAM
T94493
)
;
Two Peoples Bay Nature Reserve
, near
Picnic Area
, sifting elevated leaf litter,
34°58'27"S
,
118°10'42"E
,
1.V.2008
,
M. Rix
,
M. Harvey
,
J. Newell
,
1♀
(
WAM
T94483
)
;
Walpole-Nornalup National Park
, litter,
9.II.1979
, M.
Gray
,
1♀
(
AMS
KS14625
)
;
same data,
1♀
(
AMS
KS14636
)
;
Walpole-Nornalup National Park
,
Anderson Road
, near
Valley
of the
Giants Road
,
Tingle-Karri forest
,
leaf litter Tullgren funnel
,
34°59'32"S
,
116°52'17"E
,
3.V.2008
,
M. Rix
,
M. Harvey
, 1♁,
2♀
(
WAM
T94481
)
;
West
Cape
Howe National Park
, sifting leaf litter above
Shelley Beach
,
35°06'22"S
,
117°37'39"E
,
7.XI.2007
,
M. Rix
, 1♁,
1♀
(
WAM
T94532
)
;
West
Cape
Howe National Park
, off Torbay
Beach Road
, sifting elevated leaf litter in gully opposite
Richard Island
,
35°04'35"S
,
117°38'49"E
,
16.III.2008
,
M. Rix
,
M. Harvey
, 1♁,
2♀
(
WAM
T94531
)
;
Westcliffe
fireline track, litter,
9.II.1979
,
M. Gray
, 1♁,
1♀
(
AMS
KS14646
)
;
Witchcliffe region
,
Pentorifice Cave
(WI-122), off web near stalagmite drip at north-eastern end of cave,
25.III.1995
, R.
Foulds
, 1♁ (
WAM
T35078
)
;
near
Yate Road
, S. of
Bremer Bay
, sifting elevated leaf litter,
34°24'10"S
,
119°22'43"E
,
2.V.2008
,
M. Rix
,
M. Harvey
,
J. Newell
,
1♀
(
WAM
T94484
)
.
Etymology
.
The specific epithet is a patronym in honour J. Dale Roberts, of the University of
Western Australia
(Perth), for his exceptional supervision, patient guidance and wonderful support during M. Rix’s postgraduate study at the University of
Western Australia
.
Diagnosis
.
Males and females of
Taphiassa robertsi
can be distinguished from all other described congeners by the distinctive, bi-coloured pattern on the abdomen (
Figs 7P
, 155A–B), similar only to undescribed species in south-eastern
Australia
. In the field this species is likely to be confused only with the sympatric species
T. globosa
, which is much smaller, with relatively shorter legs.
Description
.
Holotype
male
: Total length 1.43. Carapace 0.67 long, 0.49 wide. Abdomen 0.74 long, 0.61 wide. Leg I femur 1.29. Cephalothorax, anterior sclerite, abdominal sclerotic spots brown; legs, spinneret sclerite tan-yellow; abdomen dark grey, with cream mesal stripe and cream lateral patches. Carapace raised anteriorly, fused to sternum via pleural sclerites, with circular sclerotic rim around petiole; dorsal surface of pars cephalica slightly convex in lateral view; carapace and sternum heavily punctate. Eight eyes present on anterior margin of pars cephalica; AME 0.8x diameter of ALE; PME separated by slightly less than their own diameter. Chelicerae without bulging anterior projections; promargin without peg teeth. Legs relatively long (leg I femur-carapace ratio 1.93); macrosetae absent. Abdomen globose, covered with hair-like setae, each seta projecting from small sclerotic spot; dorsal scute and lateral sclerotic strips absent. Pedipalpal patella with retrolaterally-directed, hooked lRPA and strongly recurved distal apophysis; bulb spherical, with posteriorly-bulging subtegulum; tegulum smooth, with curved ETR; embolus long (length> 5× width), curving distally (Fig. 156).
Allotype
female
: Total length 1.55. Carapace 0.68 long, 0.50 wide. Abdomen 0.82 long, 0.76 wide. Leg I femur 1.29. Cephalothorax, anterior sclerite, abdominal sclerotic spots brown; legs, spinneret sclerite tan-yellow; abdomen dark grey, with cream mesal stripe and cream lateral patches. Carapace raised anteriorly, fused to sternum via pleural sclerites, with circular sclerotic rim around petiole; dorsal surface of pars cephalica slightly convex in lateral view; carapace and sternum heavily punctate. Eight eyes present on anterior margin of pars cephalica; AME 0.8x diameter of ALE; PME separated by slightly less than their own diameter. Chelicerae without bulging anterior projections; promargin without peg teeth. Legs relatively long (leg I femur-carapace ratio 1.90); macrosetae absent. Abdomen globose, covered with hair-like setae, each seta projecting from small sclerotic spot; dorsal scute and lateral sclerotic strips absent. Pedipalp entire, five-segmented. External epigyne indistinct (Fig. 155D); spermathecae globular, almost spherical; insemination ducts short; fertilisation ducts simple, curved (Fig. 154C).
Distribution
.
Known from the western and southern coasts of south-western
Western Australia
(Fig. 215).
Remarks
.
Taphiassa robertsi
is a common and relatively widespread species in south-western
Western Australia
, where specimens have been collected by beating and sifting low vegetation and elevated leaf litter in mesic habitats. The main population extends along the south coast from Margaret River in the west to at least Bremer Bay in the east, with outlying populations in the Porongurup National Park and in caves at Yanchep National Park, north of Perth (Fig. 215). Similar species are also known from throughout south-eastern
Australia
.