The spider family Micropholcommatidae (Arachnida: Araneae: Araneoidea): a relimitation and revision at the generic level Author Rix, Michael Western Australian Museum, Welshpool DC, Perth, Australia Author Harvey, Mark Western Australian Museum, Perth, Welshpool, Australia text ZooKeys 2010 2010-02-22 36 36 1 321 journal article 10.3897/zookeys.36.306 4db6b327-7482-432e-a5f6-36f91c79fef3 1313–2970 576620 ADCACC88-6C78-4386-8E33-3F98234ECE92 Genus Pua Forster, 1959 Pua Forster, 1959: 299 . Type species by original designation Pua novaezealandiae Forster, 1959 . Brignoli, 1983: 374 . Platnick, 2009 . Affinities. The genus Pua appears to be the sister-lineage to Micropholcomma from Australia ( Fig. 4 ). Diagnosis . Pua novaezealandiae can be distinguished from all other Micropholcommatini by the presence of a short (length <5× width), hooked embolus situated distally on a rotated bulb ( Figs 29 A–C). Other diagnostic characters include the loss of the AME ( Fig. 25B ), the presence of very large, globular spermathecae (Fig. 15F), and the presence of a dorsal scute on the female abdomen ( Fig. 24B ). Description . Very small, entelegyne Araneoidea ; total length 0.80 to 1.00. Cephalothorax : Carapace without glandular depressions above maxillae ( Fig. 26D ); cuticle without glandular pits ( Fig. 26E ); margins fused to sternum via pleural sclerites. Six eyes present on anterior margin of pars cephalica ( Fig. 25B ); AME absent. Chelicerae without bulging anterior projections in males; promargin with true teeth, a pair of fused setal sockets adjacent to base of fang and single peg tooth in males ( Fig. 27F ). Legs and female pedipalp : Legs three-clawed ( Fig. 28D ), covered with smooth or serrate hair-like setae. Trichobothria present on legs; tibiae each with two (legs I–III) or three (leg IV) trichobothria ( Fig. 28C ); metatarsi without trichobothria. Female pedipalp four-segmented, with fused tibia-tarsus; claw absent ( Figs 27 C–D). Abdomen : Abdomen oval-globose; anterior sclerite present around epigastric region and petiole; large dorsal scute present on males and females ( Figs 24 A–B); lateral sclerotic strips present on males; posterior sclerotic ring surrounding spinnerets and colulus. Six spinnerets situated posterior to fleshy colulus ( Figs 23 C–D); PMS without AC gland spigots; PLS with complete triad. Anterior tracheal system well-developed, with relatively large lateral atria and multiple radiating tracheae; second tracheal tubes on each side looping mesally and extending through petiole into cephalothorax; posterior tracheal spiracle absent. Genitalia : Male pedipalp ( Fig. 29 ) relatively simple; patella with distally-directed, hooked ligulate retrolateral apophysis; bulb rotated 180 degrees, with embolus inserting distally relative to cymbium; tegulum smooth, with straight, evaginated tegular ridge; embolus exposed, short (length <5× width) and hooked; conductor absent. Female genitalia (Fig. 15F) with pair of separate, very large, globular anterior spermathecae; insemination ducts short, barely visible in dorsal view; fertilisation ducts simple, curved. Distribution . New Zealand . Composition . Only the type species Pua novaezealandiae Forster, 1959 .