A Revision of the New World Genera Brachylinga Irwin and Lyneborg and Lysilinga Irwin and Lyneborg (Diptera: Therevidae: Therevinae) with the Description of a New Genus, Elcaribe Webb Author Webb, Donald W. Author Metz, Mark A. text Zootaxa 2006 2006-08-14 1288 1288 1 241 journal article 1175­5334 FAA70D1F-49C8-40FC-9D96-CCF8C017E6BB Lysilinga nigra Webb spec. nov. ( Figs. 366 , 376, 386, 396, 406, 416, 426, 436, 444, 449) Etymology Niger (Latin) = black. This name is used as an adjective and refers to the black coloration of the specimens. Diagnosis Lysilinga nigra is taxonomically associated with L. chamela and L. digita in lacking setae on the katepisternum; in having the abdomen dark reddish brown; in having the halter knob dark brown; lacking dorsocentral macrosetae; and the hypoproct quadrate ( Fig. 366 ). Lysilinga nigra differs from L. chamela and L. digita in having the male frons setose; the anterior half of the ejaculatory apodeme cylindrical (Fig. 436); and the female abdomen dark reddish brown, glossy, pruinescence white on the posterolateral area of tergites 1–7. Description of holotype male (MEI 101362) Body length 6.8 mm . Head. Length 0.84 mm . Ommatidia smaller ventrally and laterally. Frons pruinescence silver gray, black dorsally with pair of small black areas dorsal to antennal base; setae dark reddish brown intermixed with dark yellow setae. Antenna/head length 0.69; scape and pedicel dark reddish brown, pruinescence gray, flagellum dark brown with apical third of first flagellomere dark yellow; scape length 0.22 mm , width 0.12 mm , length/width 1.8, scape/pedicel length 3.7, scape/flagellum width 0.75, setae dark reddish brown; pedicel length 0.06 mm , length/width 0.5; first flagellomere almost spherical, apical fourth attenuate apically, flagellum length 0.30 mm , width 0.20 mm , length/width 1.9, flagellum/scape length 1.4. Parafacial dark reddish brown, pruinescence gray. Maxillary palpus dark yellow; length 0.36 mm , length/width 5.1; setae dark yellow. Genal setae white. Occipital setae white becoming pale yellow, lanceolate, appressed along margin of eyes dorsally. Postocular macrosetae dark reddish brown. Thorax. Macrosetae dark reddish brown, 3 np, 2 sa, 1 pa, 0 dc, sc 1. Mesonotum dull brown; setae white, elongate intermixed with pale violet, lanceolate, appressed, polychromatic setae. Pleuron dark reddish brown, pruinescence gray; setae white on propleuron, anepisternum, laterotergite, and metanepisternum, pale violet, lanceolate, appressed, polychromatic on scutellum, absent on katepisternum. Wing. Membrane pale brown becoming darker apically; veins brown; length 4.8 mm , length/width 2.8; pterostigma brown; cell m 3 open widely. Halter dark brown. Legs. Coxal macrosetae dark reddish brown. Femora dark yellowish brown; ventral setae white on fore­ and midfemur; av 1:0:3, pv 0:0:5. Tibiae dark yellow to pale brown. Tarsi dark yellowish brown. Abdomen. Dark reddish brown to black, pruinescence white posterolaterally on tergites 5–6; dorsal setae white, appressed; lateral setae white, elongate. Terminalia (MEI 103376), brown. Tergite 8 and epandrium ( Fig. 366 ), setae dark reddish brown. Hypoproct ( Fig. 366 ) quadrate, posterior margin truncate; ending below apex of cercus; setae dark reddish brown. Gonocoxite ventral view (Fig. 376), basal three­fourths quadrate, posteromedial angle rounded, tapering apically to attenuate projection, setae dark reddish brown; dorsal view (Fig. 486) gonocoxal bridge moderately large; lateral view (Fig. 496) oblong, tapering apically to broad point; ventral lobe (Fig 376) lacking ventral spines. Gonostylus ( Fig. 406 ), basodorsal lobe absent; ventral margin straight, setae dark yellow; fine, short spines laterally. Aedeagus with dorsal apodeme (Fig. 416) square, posterolateral expansion large, pointed; ventral apodeme ( Fig. 426 ) narrow, tapered anteriorly to anterior margin of dorsal apodeme; distiphallus dorsal view (Fig. 416) elongate, tapered apically to point with dorsal carina, lateral view (Fig. 436), apical half recurved apically, subapical hood large, curved ventrally to point; ejaculatory apodeme (Fig. 416) dark yellow, anterior half narrow, cylindrical, not extending beyond anterior margin of dorsal apodeme, anterior margin rounded, in cross­section spherical. Variation (n=7). Body length 6.6–7.0, 6.8 mm . Head. Frontal setae dark reddish brown intermixed with dark yellow setae (MEI 050596, 103376). Thorax. Macrosetae 3 np, 1–2 sa, 1 pa, 0 dc, 1–2 sc. Wing. Membrane dark yellow (MEI 103376). Legs. Femora dark yellow (MEI 050596, 103376); av 1:0:3–4, pv 0:0:4–7. Female Similar to male except for following. Variation (n=10). Body length 5.8–8.0, 7.2 mm . Head. Frons black, pruinescence gray, whitish gray ventrally with pair of small, dark brown areas dorsal to antennal base; setae dark reddish brown. Antenna dark brown to black, scape and pedicel pruinescence gray. Occipital setae white, elongate becoming reddish brown, lanceolate, slightly iridescent, appressed dorsally. Thorax. Macrosetae 3 np, 2 sa, 1 pa, 0 dc, 1–2 sc. Mesonotal setae dark brown, short intermixed with white and turquoise, lanceolate, iridescent, appressed setae (on some specimens these setae appear pale violet). Legs. Femora dark yellow, dorsoapical area grading into reddish brown to dark brown; av 1–2:0:2–5, pv 0:0:2–7. Abdomen. Dark reddish brown, glossy, pruinescence white on posterolateral area of tergites 1–7; dorsal setae dark reddish brown, appressed with white, elongate, appressed setae across posterior margin of tergites 1–3; lateral setae dark reddish brown. Terminalia (MEI 101492). Anterior margin of sternite 8 slightly convex. Furca ( Fig. 444 ) oblong; length 0.35 mm , width 0.25 mm ; posteromedial lobe large cone­shaped; anterolateral and anteromedial projections absent. Common spermathecal duct ( Fig. 444 ) shorter than furca. Spermathecal duct directed anteriorly, sac­like posteriorly then becoming thread­like anteriorly. Distribution Lysilinga nigra is known from Mexico ( Fig. 449 ). Habitats and phenology Adults of Lysilinga nigra have been handnetted and collected in Malaise traps from 12 February to 3 July. Specimens examined Type specimen . The holotype male of Lysilinga nigra Webb (MEI 101362) is labeled " MEXICO , Jalisco , Careyes, 12 Feb–19 Mar 1997 , F. D. Parker" and is deposited in the USNM. PARATYPES . MEXICO . JALISCO . Careyes [Punta Careyes], 12–19.II.1997 , F. C. Parker , 1♂ 101357 ( INHS ) , 23♀ MEI 101090 ( INHS ) , 101092 (USNM), 101096, 101112, 101117–8, 101120, 101122–3, 101132, 101211, 101224, 101241, 101271–2, 101283, 101299, 101315, 101316, 101321, 101344, 101352, 101362 ( EMUS ) Malaise trap ; E stación de Biologia Chamela [19.61, ­105.05] , 3.VI.1988 , E. Ramírez , 2♀ MEI 103369, 103382 ( EBCC ) ; 12.VI.1989 , 1♀ MEI 103374 ( EBCC ) ; 13.II.1989 , 2♀ MEI 103380–1 (E BCC ) ; 20.IV.1988 , 1♀ MEI 103383 ( EBCC ) ; 11.VI.1988 , 1♀ MEI 103386 ( EBCC ) , Malaise trap . MORELOS . 2.5 km W Ajuchitlan [ 18.467 , ­98.983 ], 950 m , 15.XII.1995 , Rodriguez , A., 1♂ MEI 103379 ( MEIC ) ; 16.II.1996 , 1♀ MEI 103387 ( MEIC ) ; 16.III.1996 , 1♂ MEI 050596 ( EBCC ) ; 14.IV.1996 , 1♂ MEI 103375 ( EBCC ) ; 2.5 km N & 4 km W Huautla , Estación CEAMISH [18.45, ­99.033], 940 m , 11.IV.1996 , A. Rodriguez , 2♂ MEI 103376–7 ( EBCC ) ; 15.III.1996 , 1♀ MEI 103384 ( EBCC ) , Malaise trap . OAXACA . 8 km NW Totolapan [16.667, ­96.3], 1200 m , 3.VII.1975 , E. M. Fisher , J. L. Fisher , 1♂ MEI 147278 ( UCR ) .