Lectotypification of taxa belonging to the “ Festuca circummediterranea ” group
Author
Foggi, Bruno
Università di Firenze, Dipartimento di Biologia Evoluzionistica – Laboratori di Botanica, via G. La Pira, 4 I- 50121 Firenze, Italy.
bruno.foggi@unifiit
Author
Quercioli, Claudia
Università di Firenze, Dipartimento di Biologia Evoluzionistica – Laboratori di Botanica, via G. La Pira, 4 I- 50121 Firenze, Italy.
Author
Gennai, Matilde
Università di Firenze, Dipartimento di Biologia Evoluzionistica – Laboratori di Botanica, via G. La Pira, 4 I- 50121 Firenze, Italy.
Author
Nardi, Enio
Università di Firenze, Dipartimento di Biologia Evoluzionistica – Laboratori di Botanica, via G. La Pira, 4 I- 50121 Firenze, Italy.
Author
Signorini, Maria Adele
Dipartimento di Biotecnologie Agrarie – sez. Botanica ambientale e applicata, piazzale delle Cascine, 28 I- 50123 Firenze, Italy
text
Candollea
2012
2012-12-01
67
2
221
228
journal article
3362
10.15553/c2012v672a2
86ed7c95-806a-4447-b904-1dd97762b405
2235-3658
5718956
Festuca duriuscula
var.
campana
N. Terracc.
, Relaz. Peregr. Bot. 195. 1872
(
Fig. 2
).
Neotypus
(designated by
ALEXEEV, 1973: 106
):
ITALY
:
“
Campania
prope Sora, leg.
Terracciano
”, s.d.,
Terracciano s.n.
(
LE
[photo]!)
.
Lectotypus
(designated here for correction): with two labels:
ITALY
:
A) “
Festuca duriuscula
b. campana Terracc. [from Terracciano’s hand] /
F. ov. v. laevis subv. campana Hack
[from Hackel’s hand] / In montosis apricis aridis Campaniae; Nola/ a
Casamarciano
.
Maio 1871
/
Terracc
. [from Terracciano’s hand]” s.n.; B) “
F. laevis (Hack.) Nym. Consp.
/
ssp. laevis
/
var. heldreichii (Hack.)
/
subv. campana (Terr.) Hack.
/ III. 65. [from Markgraf-Dannenberg’s hand] det.
I. Markgraf-Dannenberg
[pr.]” (W!).
TERRACCIANO (1872: 195)
validly published a
“F.duriusculaLin. c. campana”.
The name lacks any indication of rank, but according to
MCNEILL & al. (2006
, art. 35.4), it must be regarded as a variety.
The variety is described as
“foliislongis, glaucis, culmis prope basimincrassato-tuberosis”.
It was collected by Terracciano at
“Nolasui monti di Casamarciano”,
a locality near Neaples and Caserta.
The name was typified by
ALEXEEV (1973: 106)
with a specimen housed in LE.
Fig. 2. –
Lectotypus of
Festuca duriuscula var. campana N. Terracc.
[Terracciano s.n., W] [© Naturhistorisches Museum
Wien
. Reproduced with permission]
Actually, on this lectotypification some major remarks are to be done. First of all, the specimen does not bear any note written in Nicola Terracciano’s own hand (for Terracciano’s handwriting, see
SANTANGELO & al., 1995
), neither is there any note showing that it was seen by him. Furthermore, the locality
“prope Sora”
(that is near a small town in the surroundings of Frosinone) reported on the label was not mentioned in the protologue (T ERRACCIANO, 1872). In short, the specimen cannot be regarded as strictly belonging to “original material”, as nothing shows that “the description or diagnosis validating the name was based” upon it (
MCNEILL & al., 2006
, art. 9.2, note 2). Consequently, the use of the term “
lectotype
” by A LEXEEV (1973) should be considered an “error to be corrected” (
MCNEILL & al., 2006
, art. 9.8), as the specimen selected by him is not actually a
lectotype
, but a
neotype
.
Furthermore, it must also be noted that the specimen does not fit the original description, as it lacks the diagnostic character
“culmis prope basim incrassato-tuberosis”
(i.e. “culms swollen like tubers near the base”). As a matter of fact, in the specimen the culms are entirely slender, even to the base.
Searching for original Terracciano’s material, in NAP we could find
two specimens
collected at “Casamarciano”, the locality reported in the protologue. Both specimens bear labels with notes in Nicola Terracciano’s own hand and perfectly fit the original description of
F.duriuscula
var.
campana
. Unfortunately, neither of them shows labels or notes with the epithet “campana”, so they may not be considered as
type
material.
In W
, we found one more specimen from Hackel’s herbarium collected at Casamarciano, bearing a label handwritten by Terracciano where the full name of the new plant is reported:
“F.duriusculab.campana”
. We do not know how this specimen arrived in
Wien
: possibly it was sent to Hackel as a gift by Terracciano himself, or by a curator of NAP. It comes from the
locus classicus
and perfectly fits the description reported in the protologue, with the character
“culmisprope basimincrassato-tuberosis”
particularly evident. For these reasons, we propose to supersede Alexeev’s neotypification according to
MCNEILL & al. (2006
, art. 9.11, art. 9.17) and we designate this specimen as the
lectotype
of the name.
This taxon was raised to the rank of species,
F.campana
(N. Terracc.) Alexeev, by
ALEXEEV (1973: 105-106)
.
Even at the rank of species, the name retains the same
type
designed by us for
F. duriuscula
var.
campana
.
It can be added that a close analysis on this specimen showed that the swollen culms described in the protologue are actually galls due to an insect (
Tetramesa
cfr.
brevicormis
,
Hymenoptera
), which uses to lay eggs inside the stems of this plants.