Taxonomic study of the genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) X. The commotica species-group, with descriptions of twenty-two new species Author Wang, Shuxia 0000-0002-9316-6661 College of Life Sciences, Nankai University, Tianjin 300071, China & shxwang @ nankai. edu. cn; https: // orcid. org / 0000 - 0002 - 9316 - 6661 shxwang@nankai.edu.cn Author Liu, Chen 0000-0002-9111-6644 College of Life Sciences, Nankai University, Tianjin 300071, China & 348143302 @ qq. com; https: // orcid. org / 0000 - 0002 - 9111 - 6644 348143302@qq.com text Zootaxa 2021 2021-06-01 4980 2 293 330 journal article 6009 10.11646/zootaxa.4980.2.4 88b4aa3d-deff-4fb9-a9eb-1aa9e2726140 1175-5326 4889068 6E352CB3-CF9B-4EE8-B17C-67C88B35FE47 Promalactis denticuligera Wang , sp. nov. ( Figs 10 , 32 , 49 ) Type material. CHINA , Tibet : Holotype ♂ , Tongmai Town ( 30.10°N , 95.08°E ), 2029 m , 12.VIII.2018 , leg. MJ Qi , slide No. LC 19310 . Paratypes ( 23♂ , 55♀ ): 6♂ , 39♀ , same data as holotype, except 12‒16.VIII.2019 , slide No. LC 19311 ♀ ; 14♂ , 5♀ , same data as holotype, except 23‒26. VI .2009 , leg. MJ Qi & JQ Deng , slide Nos. LC 19312 ♂ , LC19313 ♀ ; 2♀ , Yadong Village , Motuo County , 833 m , 16.VIII.2017 , leg. XF Yang & MJ Qi , slide No. JYY17327 ; 1♂ , 9♀ , Pailong Town , Linzhi County , 2031 m , 17‒18.VIII.2018 , leg. MJ Qi , slide Nos. LC 19326 ♂ , LC19331 ♀ ; 1♂ , Pailong Town , Linzhi County , 2010 m , 6.VII.2013 , leg. JW Li , slide No. LC 19353 ; 1♂ , Linzhi Town , Linzhi City , 2985 m , 19. VI .2019 , leg. MJ Qi & JQ Deng , slide No. LC 19316 . Diagnosis. The new species can be distinguished from other members of the group by two features of the male genitalia: the asymmetrically bilobed valvae, and the costa of the left valva produced into a free dorsodistal lobe bearing a cluster of spines along the costal margin. Description . Adult ( Fig. 10 ). Forewing length 7.0‒9.0 mm. Head: Frons greyish brown, vertex cream white, occiput rust brown. Labial palpus with second palpomere rust brown on outer surface, ochreous yellow on inner surface; third palpomere black on outer surface, rust brown on inner surface except white apically. Antenna with scape white dorsally, pale grey ventrally; flagellum black annulated with white on dorsal surface, black on ventral surface. Thorax: Dorsum of mesothorax and tegula rust brown. Forewing ochreous brown on basal 2/3, ochreous yellow on distal 1/3; markings white edged with black scales: costal spot triangular, from distal 1/3 narrowed to above posterior angle of cell; basal streak slender, straight, oblique inward to base of fold; antemedian streak from dorsal 1/3 to anterior margin of cell, parallel with basal streak; dorsal streak from basal 2/3 extending obliquely outward to distal 1/4 of posterior margin of cell; tornal spot black, narrowly diffused inward to end of dorsal streak; distal patch black, extending from apex along termen to beyond tornal spot, gradually narrowed; apical spot sub-oval, small, set in black distal patch; fringe ochreous yellow except grey before tornus. Hindwing and fringe grey. Fore- and midlegs black, except fore femur white and midleg yellowish brown on ventral surface, tibiae with white spot at base and middle, with a tuft of white scales at apex, basal tarsomere white at base and apex, second tarsomere white apically; hindleg yellowish brown, tibia with long grey hairs, basal tarsomere white apically, second and third tarsomeres black except white distally, fourth and fifth tarsomeres white. Abdomen: Male genitalia ( Fig. 32 ). Uncus wide parallel-sided from base to about basal 2/5, then slightly narrowed to rounded apex. Gnathos longer than uncus; mesial plate triangular, wide at base, narrowed to pointed apex, granulate distally; basal arms wide basally, narrowed to mesial plate. Tegumen with height more than three times length of uncus; lateral arms widened and arched on inner margin near apex. Valvae narrow at base, widened toward apex, asymmetrically bilobed distally; left valva: dorsal lobe slender, slightly narrowed to apex, ventral lobe wide at base, narrowed to basal 2/3, then slender to rounded apex, setose, shorter than dorsal lobe; costa very narrow, with clustered spines along costal margin in distal half; sacculus broadly banded, almost uniform in width, with a large semicircular concavity dorsally before free distal process; right valva: costa narrow, produced to a free digitate dorsal lobe distally, ventral lobe much larger and longer than dorsa lobe, elongate sub-triangular, with a flat apex; sacculus narrowly banded, wide at base, narrowed to basal 2/3, fused with valva and setose in distal 1/3. Saccus longer than uncus, elongate triangular, wide at base, narrowed to rounded apex. Juxta a large sclerotized plate, reaching posterior 1/4 of tegumen, narrow basally, widened distally, rounded apically; basal lobe clubbed, reaching middle of saccus. Aedeagus straight, almost as long as left valva, tubular, distal 1/3 spatulate, denticulate laterally and apically. Cornutus absent. Female genitalia ( Fig. 49 ). Apophyses posteriores about 1/3 length of apophyses anteriores. Eighth sternum sub-rectangular, with a notch at middle on posterior margin.Antrum heavily sclerotized, slightly narrowed to ductus bursae. Ductus bursae membranous, slightly inflated, less than half length of antrum, with a coiled belt posteriorly. Corpus bursae slightly longer than ductus bursae, membranous, ovate; signum sub-ovate, with dense, large and small denticles. Distribution . China ( Tibet ). Etymology. The specific epithet is derived from the Latin denticuliger , referring to the distally and apically toothed aedeagus.