Taxonomic study of the genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) X. The commotica species-group, with descriptions of twenty-two new species Author Wang, Shuxia 0000-0002-9316-6661 College of Life Sciences, Nankai University, Tianjin 300071, China & shxwang @ nankai. edu. cn; https: // orcid. org / 0000 - 0002 - 9316 - 6661 shxwang@nankai.edu.cn Author Liu, Chen 0000-0002-9111-6644 College of Life Sciences, Nankai University, Tianjin 300071, China & 348143302 @ qq. com; https: // orcid. org / 0000 - 0002 - 9111 - 6644 348143302@qq.com text Zootaxa 2021 2021-06-01 4980 2 293 330 journal article 6009 10.11646/zootaxa.4980.2.4 88b4aa3d-deff-4fb9-a9eb-1aa9e2726140 1175-5326 4889068 6E352CB3-CF9B-4EE8-B17C-67C88B35FE47 Promalactis bicornuta Wang , sp. nov. ( Figs 5 , 27 ) Type material. CHINA , Guizhou : Holotype ♂ , Xiasi Village , Kuankuoshui Reserves , Suiyang County , 840 m , 9.VI.2010 , leg. LL Yang , slide No. DZH10246. Diagnosis. The new species can be distinguished from other members of the group by the large, sub-rectangular, preapical process that extends from the dorsal margin of the saccular lobe of the valva to the ventral margin of the costal lobe, and the aedeagus with a distinct apical spine-shaped process. Description . Adult ( Fig. 5 ). Forewing length 4.5 mm . Head: Vertex bright leaden, occiput mottled with silvery grey. Labial palpus with first and second palpomeres dark grey on inner surface, dark brown mottled with black scales on outer surface; third palpomere black except white at base and apex. Antenna with scape white, black apically on dorsal surface, pecten dark brown; flagellum black, with white annulation on dorsal surface, dark brown on ventral surface. Thorax: Dorsum of mesothorax and tegula ocherous brown. Forewing yellowish brown, costal margin black on basal half; markings white edged with black scales: costal spot at basal 3/5, rounded, with dense black scales anterolaterally; cell with a small spot at outer margin below costal spot; basal streak extending to base of fold; antemedian streak falcate, from dorsal 1/3 extending outward to basal 1/3 of anterior margin of cell, arched inward to basal 1/5 of cell, forming a right angle; dorsal streak from middle of dorsum extending to distal 1/4 of posterior margin of cell, continued to tornus by a black line; pretornal spot represented with a few white scales; both apical and terminal spots elliptic, encircled by dense black scales; fringe ochreous yellow except black at base of termen. Hindwing and fringe deep grey. Fore- and midlegs black, except coxa yellow, tibia with a white spot at base and middle respectively, with a tuft of white scales at apex, tarsus white at apex of basal tarsomere and on entire second tarsomere; midleg grey ventrally, black dorsally, tibia with a white spot at base, with a tuft of white scales at middle and apex, tarsus white at apices of basal two tarsomeres; hindleg yellow ventrally, black dorsally, except tarsus white at apices of basal four tarsomeres and on entire apical trasomere. Abdomen: Male genitalia ( Fig. 27 ). Uncus bell-shaped, broad at base, narrowed to basal 2/3, distal 1/3 slender, digitate, rounded apically. Gnathos slightly shorter than uncus; mesial plate lingulate, rounded at apex, with a membranous fine apical process, squamous in distal 1/3; basal arms approximately 1/3 length of mesial plate. Tegumen with lateral arm narrowed anteriorly, obtuse apically. Valva sub-rectangular, bilobed apically: costa slightly concave basally, produced to a apex-rounded costal lobe slightly bent ventrad distally; sacculus narrow, produced to a setose, apex-rounded saccular lobe distally, large sub-rectangular preapical process extending from its dorsal margin to ventral margin of costal lobe, with a short spine on outer margin apically. Saccus approximately 2.3 times length of uncus, wide at base, narrowed to basal 1/3, distal 2/3 slender, rounded apically. Juxta sub-elliptical, weakly sclerotized, laterally with a spine-like process at basal 1/3; basal lobe short, clubbed. Aedeagus straight, approximately 1.5 times length of valva, with a long spine-shaped process apically; with two cornuti: one cornutus slender, approximately 1/4 length of aedeagus, located medially, the other stout, approximately 1/7 length aedeagus, located distally. Female. Unknown. Distribution: China ( Guizhou ). Etymology. The specific epithet of the new species is derived from the Latin prefix bi- and cornutus , referring to the two cornuti of the aedeagus in the male genitalia.