New taxa of Tateidae (Caenogastropoda, Truncatelloidea) from springs associated with the Great Artesian Basin and Einasleigh Uplands, Queensland, with the description of two related taxa from eastern coastal drainages Author Zhang, - H. text Zootaxa 2019 2019-04-10 4583 1 1 67 journal article 27316 10.11646/zootaxa.4583.1.1 027ed27d-5035-4646-93a6-c23b565f3d20 1175-5326 2637603 27F24995-359E-46F6-AB22-75568BACFDCF Edgbastonia ( Barcaldinia ) rugosa n. sp. Material examined. Holotype : Queensland , Myross Station , small spring, 22° 46' 32" S , 145° 26' 17" E , under two large gum trees, W.F. Ponder , R. Fairfax & R. Fensham , 17 Sep 2006 , C.479949 . Paratypes : Same data, C.457753, 20+; QM MO85768 , 5 . Other material examined from the same locality : C.483133, 15; C.483138, 3; C.483134, 6; C.483137, 1. Other material examined: Queensland , Myross Station , 22° 47' 54" S , 145° 26' 39" E , R.J. Fensham & R. Fairfax , 1 Jul 2008 , C.483135, 1 subadult . Shell ( Fig. 13F ). Elongate-conic, spire outline moderately convex, normally coiled, translucent. Length 2.0– 2.7 mm (mean 2.4 mm ), width 1.0– 1.2 mm (mean 1.1 mm ). Protoconch about 1.3 whorls, minutely pitted. Teleoconch whorls weakly convex, evenly rounded, total number 5.2–5.7 (mean 5.5). Sculptured with weak, rounded axial ribs. Umbilicus represented by chink only. Aperture ovate or pyriform, inner lip narrow, thin or medium, slightly separated along whole length of parietal wall, outer lip thin or medium. Periostracum moderately developed, white or reddish-brown. Operculum ( Fig. 14H ). Translucent, yellow-brown, slightly concave. Inner side simple, with white streak or smear present, protuberance lacking. Head-foot and external body. Snout, tentacles, neck, dorsal and lateral foot and opercular lobes pigmented, mantle roof black, visceral coil densely pigmented. Mantle cavity. Ctenidium well-developed, filaments 21–27, broadly triangular, apex right edge. Osphradium narrowly oval, towards posterior end of ctenidium, length relative to gill 0.25–0.32. Hypobranchial gland moderately developed, or thin (poorly developed). Rectum with U-shaped bend, anus behind mantle collar. Kidney half or more in roof of mantle cavity. Renal gland transverse. Pericardium half or more in roof of mantle cavity, with short separation from posterior end of ctenidium. TABLE 13. Shell measurements (in mm) of type material of Edgbastonia ( Barcaldinia ) rugosa n. sp. (paratypes, C.457753). See Materials and Methods for abbreviations.

SH	SW	AH	AW	BW	TW
Holotype	2.55	1.12	0.87	0.60	1.57	5
Paratypes (5)
Min	2.01	1.00	0.72	0.65	1.34	5.20
Max	2.68	1.19	0.85	0.84	1.58	5.70
Mean	2.42	1.09	0.77	0.75	1.48	5.46
SD	0.26	0.07	0.05	0.07	0.09	0.23

Radula ( Fig. 19A, B ). Central teeth with cusp formula 5–6+1+5–6, basal cusps 1–2+1–2; median cusp sharp, about 1.5 times length of adjacent cusps. Lateral teeth with cusp formula 4–5+1+4–6; main cusp sharp, about 1.5 times length of adjacent cusps. Inner marginal teeth with 23–28 cusps. Outer marginal teeth with 24–27 cusps. Female reproductive system ( Fig. 16I , L ). Ovary simple sac. Renal oviduct with short, U-shaped, vertical arch. Seminal receptacle near anterior edge of bursa, oval, duct very short; traces of orientated sperm also in renal oviduct and/or bursal duct. Bursa copulatrix behind albumen gland, triangular, shorter than albumen gland, bursal duct enters bursa mid anteriorly or antero-ventrally, bursal duct joins coiled oviduct at posterior mantle cavity wall. Albumen gland behind posterior mantle cavity wall. Capsule gland with two distinct glandular zones, medium thickness in cross section, markedly indented by rectum. Anterior vestibule small, opening subterminal, long, cowl and/or gutter associated with oviduct opening absent. Male reproductive system ( Fig. 17E , 18F ). Prostate gland less than half in mantle roof, pyriform, thick in cross section. Posterior pallial vas deferens slightly undulating or coiled, anteriorly strongly undulating. Penis towards middle of head, well down neck, distal end tapering, simple, terminal papilla absent. Etymology. Rugosa—Latin, wrinkled. Distribution and habitat. Known mainly from one small shallow spring on Myross Station ( Fig. 1 ) and one other nearby location. Lives sympatrically with Ed . ( B .) pagoda n. sp. and Ed. ( B .) cf. pallida . Remarks. This species is most similar to Ed. ( B .) acuminata in shell morphology, having weakly convex whorls, but differs in having fewer whorls and a shorter shell, and in being sculptured with weak, rounded axial ribs. This latter character is unique in the Jardinella group although narrow, rather sharp ribs are seen in Ed . ( B .) corrugata corrugata . Anatomically, this species is closest to Ed. ( B .) acuminata and Ed. ( B .) zeidlerorum . A discriminant function analysis using five shell measurements was carried out to compare all four of the tallspired species from the Edgbaston and Myross springs ( Fig. 20C ). Overall 85% of the specimens were correctly identified (Wilks’s Lambda 0.035 p<0.000). All specimens of Ed. zeidlerorum , Ed. rugosa and Ed. acuminata were correctly identified. In contrast, 73% of Ed. jesswiseae were correctly identified (with all the misidentified specimens being allocated to Ed. pagoda ) and 70% of Ed. pagoda (with all the misidentified specimens being allocated to Ed. jesswiseae ).