Hermit crabs of the genera Calcinus Dana, Clibanarius Dana, and Dardanus Paul’son from the PANGLAO 2004 Expedition, with description of a new species and a checklist of the hermit crabs of the Philippines (Crustacea: Anomura: Paguroidea) Author Malay, Maria Celia (Machel) D. Author Rahayu, Dwi Listyo Author Chan, Tin-Yam text Raffles Bulletin of Zoology 2018 2018-01-17 66 23 65 journal article 6557 10.5281/zenodo.4503028 479d8c3b-8194-4f81-bed5-d157d5aa021e 2345-7600 4503028 C0EDB2F2-78E4-43A0-A8C4-E4165DF6406F Calcinus latens ( Randall, 1840 ) ( Fig. 1E ) Pagurus latens Randall, 1840: 135–136 ( Hawai’i , type locality). Calcinus terrae-reginae – Estampador, 1937: 505 ; Yap-Chiongco, 1938: 206–207 ; Estampador, 1959: 53 . Calcinus latens – Grant & McCulloch, 1906: 34–35 ; Estampador, 1937: 505 ; Yap-Chiongco, 1938: 207–208 ; Fize & Serène, 1955: 58–66 , Fig. 9 , pl. 2 nos. 9–11; Estampador, 1959: 53 ; Ball & Haig, 1972: 101–102 ; Wooster, 1984: 154–156 ; Haig & Ball, 1988: 160 ; Morgan, 1991: 890–983, Figs. 34–36; Gherardi & McLaughlin, 1994: 622–624 ; Poupin, 1997: 703–704 , Figs. 3H , 6C ; Hoover, 1998: 256 , 1 unlabelled figure; McLaughlin et al., 2007: 167–169, 2 unlabelled figures; Malay & Paulay, 2010 : Appendix S1. Calcinus abrolhensis Morgan, 1988: 218–222 , Fig. 1 (Houtman Abrolhos, Australia , type locality). Material examined. 1 male SL 2.8 mm ( ZRC ), stn D5, Pontod Islet , Panglao I. , 9°33.6′N , 123°43.5′E , 0–3 m , 6.vi.2004 ; 2 males SL 1.8–3.9 mm , 1 indiv. in shell ( ZRC ), stn D8, Dao , Panglao I. , 9°34.0′N , 123°48.9′E , 1–4 m , 9.vi.2004 ; 1 female SL 2.4 mm , 2 males SL 0.8–1.4 mm , 3 indivs. in shell ( ZRC ), stn M1, Alona Beach , Panglao I. , 9°32.9′N , 123°46.6′E , 0–1 m , v.2004 to vii.2004 ; 4 males SL 3.8–4.9 mm ( ZRC ), stn M2, west end of Alona Beach , Panglao I. , 9°32.8′N , 123°45.9′E , 0–2 m , 30.v.2004 ; 1 female SL 3.1 mm , 3 males SL 3.1–3.5 mm ( ZRC ), stn M3, Danao , Panglao I. , 9°32.5′/33.1′N, 123°44.7′/45.5′E, 0–2.5 m , 31.v.2004 ; 1 male SL 5.5 mm ( ZRC ), stn M5, Doljo Point , Panglao I. , 9°35.5′N , 123°43.3′/44.3′E , 0–2 m , v.2004 to vi.2004 ; 1 female SL 2.5 mm , 1 male SL 5.0 mm ( ZRC ), stn M7, Momo Beach , Panglao I. , 9°36.1′N , 123°45.2′E , 0–3 m , 1.vi.2004 ; 2 females SL 4.5–4.9 mm , 2 males SL 3.6–5.9 mm ( ZRC ), stn M18, Gak-Ang Islet , off Panglao I. , 9°33.0′N , 123°45.5′E , 0–1 m , 10 & 12.vi.2004 ; 1 female SL 2.5 mm , 1 male SL 3.1 mm ( ZRC ), stn M20, Pamilacan I., 9°29.3′N , 123°55.1′E , 0–2 m , 13.vi.2004 ; 29 indivs. not sexed and measured ( ZRC ), stn M25, Danao , Panglao I. , 9°33.1′N , 123°44.6′E , 0–1 m , 17.vi.2004 ; 1 ovig. female SL 4.2 mm , 14 males SL 2.4–5.2 mm , 8 indivs. in shell ( ZRC ), stn M51, Mayacabac , Panglao I. , 9°36.8′N , 123°52.2′E , 0 m, 30.vi.2004 ; 3 males SL 1.3–2.9 mm ( ZRC ), stn M58, Balicasag I., 9°31.3′N , 123°41.0′E , depth unknown, 4.vii.2004 ; 1 male SL 3.0 mm, stn R1, Balicasag I., 9°31.2′N , 123°41.3′E , 5–7 m , 30.v.2004 ; 1 male SL 2.6 mm ( ZRC ), stn R10, Biking , Panglao I. , 9°35.3′N , 123°50.5′E , 2–10 m , 1.vi.2004 ; 1 female 2.0 mm ( ZRC ), stn R11, Biking , Panglao I. , 9°35.2′N , 123°50.5′E , 4 m , 2.vi.2004 ; 1 male SL 3.9 mm ( ZRC ), stn R24, Bingag , Panglao I. , 9°37.5′N , 123°46.8′E , 0–2 m , 6.vi.2004 ; 2 females SL 3.6–4.5 mm ( ZRC ), stn R36, Pamilacan I., 9°30.2′N , 123°55.3′E , 3–32 m , 9.vi.2004 ; 3 males SL 2.7–5.4 mm ( ZRC ), stn R38, Pamilacan I., 9°29.4′N , 123°56.0′E , 6–37 m , 11.vi.2004 ; 1 male SL 5.4 mm ( ZRC ), stn R44, lagoon near Doljo Point , Panglao I. , 9°33.3′/34.6′N, 123°43.9′/43.4′E, 2 m , 14.vi.2004 ; 1 male SL 4.5 mm ( ZRC ), stn R50, Looc , Panglao I. , 9°35.7′N , 123°44.4′E , 3–7 m , 17.vi.2004 ; 1 male SL 3.4 mm ( ZRC ), stn R53, Sungcolan Bay , Panglao I. , 9°38.4′N , 123°49.7′E , 3–5 m , 20.vi.2004 ; 1 ovig. female SL 4.5 mm ( ZRC ), stn R58, Looc (lagoon side), Panglao I. , 1–3 m , 9°35.7′N , 123°44.7′E , 1–3 m , 22.vi.2004 ; 1 indiv. in shell ( ZRC ), stn R66, Sungcolan inlet, Panglao I. , 9°38.3′N , 123°50.3′E , 1–3 m , 28.vi.2004 . Habitat. Intertidal to subtidal, 0 to approximately 10 m deep, reef flats, seagrass beds, and reef slopes, on sandy, sand-mud, rocky, coralline, and soft bottoms. Remarks. Yap-Chiongco (1938) and Estampador (1937 , 1959 ) report both C. latens and C. terrae-reginae from the Philippine waters. Calcinus terrae-reginae was synonymised with C. latens by Grant & McCulloch (1906) . Calcinus latens is apparently a species complex, with three distinct forms found in Hawai’i, the Gulf of Oman , and the wider Indo-West Pacific region ( Malay & Paulay, 2010 ). This species complex is very abundant throughout its range, and occupies a wide variety of different habitats.