Darwin wasps of the subfamily Pimplinae (Hymenoptera: Ichneumonidae) of Mexico: Ephialtes genus-group Author Khalaim, Andrey I. Facultad de Ingeniería y Ciencias, Universidad Autónoma de Tamaulipas, Cd. Victoria, Tamaulipas, Mexico. & Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia. Author Ruíz-Cancino, Enrique 0000-0003-4305-6568 Facultad de Ingeniería y Ciencias, Universidad Autónoma de Tamaulipas, Cd. Victoria, Tamaulipas, Mexico. & https: // orcid. org / 0000 - 0003 - 4305 - 6568 text Zootaxa 2022 2022-07-29 5169 3 201 251 journal article 112030 10.11646/zootaxa.5169.3.1 2cbc0f84-2011-4e7c-868f-6c305d5f3308 1175-5326 6952172 BB6FBD07-6B8B-4615-BD66-C4AEB612145F 1. Anastelgis chiapasana Khalaim & Ruíz-Cancino , sp. nov. Material examined. Holotype female ( EMEC ), Mexico , Chiapas , Lagunas de Montebello , Sumidero de Río Comitán , 1372 m , 31.viii.1974 , coll. D.E. & J.A. Breedlove. Description. Female . Fore wing length about 8.0 mm. Upper tooth of mandible blunt, slightly longer and much broader than the lower tooth. Malar space very short, about 0.15× as long as basal mandibular width. Face somewhat transverse, 1.2× as broad as long, slightly convex, polished, with scattered punctures. Genae short, strongly rounded behind eyes in dorsal view. Lateral ocellus separated from eye by one its own maximum diameter. Occipital carina fine, complete, joining hypostomal carina far above base of mandible. Epomia strong, as subvertical slightly curved ridge. Mesoscutum smooth and shining, with fine punctures and sparse setae. Notaulus distinctly impressed anterolaterally. Mesopleuron polished, very finely and very sparsely punctate. Epicnemial carina reaching above the level of lower corner of pronotum. Metapleuron convex, polished. Submetapleural carina strong, complete, in lateral view clearly visible for its entire length. Propodeum in profile moderately convex, with lateromedian longitudinal carinae discernible as very small tubercles anteriorly, otherwise dorsally smooth. Pleural carina complete. Fore wing with vein 1cu-a opposite M&RS. Hind wing with nervellus intercepted slightly below its middle. Metasoma long and slender. First tergite broadened posteriorly, 1.7× as long as posteriorly broad, dorsally mostly smooth; first tergite dorsally concave in anterior part, the concavity bounded laterally by strong and posteriorly convergent carinae which are distinct in anterior 0.3 of tergite and obsolescent centrally (as in Fig. 6 ). Upper margin of first tergite, in profile, slightly concave anteriorly and weakly convex posteriorly. Second tergite 1.35× as long as maximally broad, with flattened triangular dull areas anterolaterally, centrally distinctly and densely punctate (distance between punctures equal to or slightly shorter than diameter of one puncture). Third tergite 1.25× as long as maximally broad, with a pair of nearly smooth lateromedian rounded swellings. Tergites 4 and 5 similar, with punctures sparser and weaker. Subgenital plate with broad V-shaped membranous area anteriorly. Ovipositor decurved, projecting beyond apex of metasoma by 4.4× length of hind tibia; sheath with long and dense pubescense. Head black, clypeus and mandible (teeth black) dark brownish black, face with slight brown tinge, palpi white. Antenna with scape and pedicel pale ventrally and blackish dorsally, flagellum entirely black. Mesosoma immaculately orange-brown with only propleuron ventrally (next to fore coxa) pale and extreme hind margin of propodeum black; tegula white in anterior half and brown in posterior half. Pterostigma brown. Legs predominantly orange-brown; fore and mid coxae, trochanters and trochantelli white; hind tarsomere 5 infuscate. Metasoma orangebrown, tergites 2–5 with posterolateral black marks. Ovipositor sheath black with apical 0.2 white. Male . Unknown. Etymology. The new species is named after the type locality, [State of] Chiapas . Distribution. Mexico ( Chiapas ). Comparison. In the key to the Costa Rican species of Anastelgis ( Gauld et al . 1998: 22 ) , the new species runs to couplet 4, but does not match neither A. garciai Gauld nor A. jaramilloi Gauld in that couplet as it has mesosoma entirely orange-brown, second tergite distinctly and closely punctate centrally, upper tooth of mandibe blunt and much broader than the lower tooth, and epomia quite strong. Anastelgis chiapasana sp. nov. possesses genae, in front view, almost straight, similar to those in A. jaramilloi (see Fig. 261 in Gauld 1991: 199 ), not convex as in A. garciai . The new species also differs from Central American A. impossibilita Gauld et al . by its orange-brown legs with only fore and mid coxae, trochanters and trochantelli white, and from the Nearctic species A. terminalis Townes by its orange-brown mesosoma without black markings, and longer metasomal tergites 2 and 3.