Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae) Author Shaverdo, Helena Author Sagata, Katayo Author Balke, Michael text ZooKeys 2018 803 7 70 http://dx.doi.org/10.3897/zookeys.803.28903 journal article http://dx.doi.org/10.3897/zookeys.803.28903 1313-2970-803-7 936CFD88F297440EA9BE4C258AE9BD09 6. Exocelina desii (Balke, 1999) Figs 15, 39 Copelatus (Papuadytes) desii Balke, 1999: 274; Nilsson 2001 : 76 (catalogue). Papuadytes desii (Balke, 1999): Nilsson and Fery 2006 : 56 (comb. n.). Exocelina desii (Balke, 1999): Nilsson 2007 : 33 (comb. n.); Nilsson and Hajek 2018 : 65 (catalogue). Exocelina desii MB1399: Toussaint et al. 2014 : supplementary figs 1-4, table 2; Toussaint et al. 2015 : supplementary figs S1-S2, table S3, and information S5-S6. Type locality. Papua New Guinea: East Sepik Province, Amboin Patrol Post, Karawari Lodge, ca 04°29'05.8"S , 143°26'37.5"E , <100 m a.s.l. Type material studied. Paratype: female "Papau [sic!] New Guinea East Sepik Province, Amboin Patrol Post, Karawari Lodge 14Jan.1983, A.C.Messer", "Paratypus Copelatus desii M. Balke des. 1999" [red] (NHMW). Note: The holotype has not been found. According to Balke (1999) , it was deposited in the Natural Museum of Natural History, Smithsonian Institution, Washington, DC, USA. Additional material. Western Highlands: 3 females "Papua New Guinea: Western Highlands, Mt. Hagen town area, 1600m, 7.xii.1994 05.49.745S 144.22.357E Balke & Kinibel (PNG 131)" (ZSM). 1 male, 1 female "Papua New Guinea: Western Highlands, Kurumul, 6Km SW Kudjip, small stream, 1584m, 13.vi.1994, 05.53.426S 144.36.600E, John (PNG 78)", the female with an additional green label "M.Balke 1342" (ZSM). Eastern Highlands: 5 males "Papua New Guinea: Eastern Highlands, Akameku-Brahmin, Bismarck Range, 1900m, 23.xi.1994, 05.54.284S 145.22.271E, Balke & Kinibel (PNG 108)", one of them with an additional green label "M.Balke 1399" (NHMW, ZSM). 3 males, 2 females "Papua New Guinea: Eastern Highlands, Akameku-Brahmin, Bismarck Range, 1500m, 24.xi.1994, 05.51.964S 145.23.604E, Balke & Kinibel (PNG 111)" (NHMW, ZSM). Simbu: 3 males, 1 female "Collection Naturhistorisches Museum Basel", "Papua New Guinea Simbu prov. L. Ciek lgt.", Kundiawa, Mu vill. 145°02'E 4°42'S III.2001, 1900m" (NHMB). 1 male "Ibisca Niugini, PNG 28-30.x.2012 Mount Wilhelm 1700m", " -5,759269238 145,235611 FIT-MW1700-K-2/8-d01 / Plot 11 / P1959 Vial 05833" (IECA). 1 male " -5,759269238 145,235611 FIT-MW1700-J-1/8-d01 / Plot 10 / P1950 Vial 02485", "Ibisca Niugini, PNG 25-27.x.2012 Mount Wilhelm 1700m" (ZSM). 2 females "Ibisca Niugini, PNG 27-29.x.2012 Mount Wilhelm 1700m -5,759269238 145,235611 MW1700 / P1943 Vial 04017" (ZSM). 1 female "Ibisca Niugini, PNG 31.x-2.xi.2012 Mount Wilhelm 1700m -5,759269238 145,235611 MW1700 / P1953 Vial 07522" (ZSM). 1 female "Ibisca Niugini, PNG 2-4.xi.2012 Mount Wilhelm 1700m -5,759269238 145,235611 MW1700 / P1914 Vial 07577" (ZSM). 1 female "Ibisca Niugini, PNG 25-27.x.2012 Mount Wilhelm 1700m -5,759269238 145,235611 MW1700 / P1878 Vial 02411" (ZSM). 1 female "Ibisca Niugini, PNG 5-7.xi.2012 Mount Wilhelm 1700m", " -5,759269238 145,235611 FIT-MW1700-O-6/8-d12 / Plot 15 / P1995 Vial 05666" (ZSM). 1 female "Ibisca Niugini, PNG 27-29.x.2012 Mount Wilhelm 1700m -5,759269238 145,235611 ", "FIT-MW1700-H-2/8-d03 / Plot 8 / P1935 Vial P1935-CODYTI" (ZSM). 1 female "Ibisca Niugini, PNG 27-29.x.2012 Mount Wilhelm 1700m -5,759269238 145,235611 ", "FIT-MW1700-I-2/8-d03 / Plot 9 / P1943 Vial P1943-CODYTI" (ZSM). 1 female "Ibisca Niugini, PNG 28-30.x.2012 Mount Wilhelm 1700m", " -5,759269238 145,235611 FIT-MW1700-T-2/8-d04 / Plot 20 / P2031 Vial 05553" (ZSM). 1 female "Ibisca Niugini, PNG 27-29.x.2012 Mount Wilhelm 1700m", " -5,760916233 145,2353363 MW1700 / P1895 Vial P1895-CODYTI" (ZSM). 1 female "Ibisca Niugini, PNG 4-6.xi.2012 Mount Wilhelm 1700m", " -5,760916233 145,2353363 FIT-MW1700-C-6/8-d11 / Plot 3 / P1899 Vial 04070" (ZSM). 1 female "Ibisca Niugini, PNG 25-27.x.2012 Mount Wilhelm 1700m -5,759910107 145,234726 MW1700 / P1886 Vial 07540" (ZSM). 1 female "Ibisca Niugini, PNG 28-30.x.2012 Mount Wilhelm 2200m", " -5,75897789 145,1860657 FIT-MW2200-F-7/8-d13 / Plot 6 / P2314 Vial 07373-CODYTI" (ZSM). 1 female "Ibisca Niugini, PNG 27-29.x.2012 Mount Wilhelm 2200m", " -5,75897789 145,1860657 FIT-MW2200-K-6/8-d12 / Plot 11 / P2353 Vial 11687" (ZSM). 1 female "Ibisca Niugini, PNG 28-30.x.2012 Mount Wilhelm 2200m -5,760178089 145,186264 MW2200 / P2282 Vial 07152" (ZSM). 2 females "Ibisca Niugini, PNG 29-31.x.2012 Mount Wilhelm 2200m -5,760178089 145,186264 FIT-MW2200-N-7/8-d14/ Plot 14 / P2378 Vial 15681" (ZSM). 2 females "Ibisca Niugini, PNG 27-29.x.2012 Mount Wilhelm 2200m FIT-MW2200-O-6/8-d12 / Plot 15 / P2385 Vial ...-CODYTI" (ZSM). 1 female "04103 Ibisca Niugini 2012" (ZSM). 1 female "04103" (ZSM). Diagnosis. For complete description, see Balke (1999 : 274). Beetle medium-sized: TL-H 3.5-4.3 mm, oblong-oval; reddish brown to brown, with paler pronotum; submatt, with dense, rather coarse punctation and rather strongly impressed microreticulation; pronotum with distinct lateral bead; male antennae simple (Fig. 15); male protarsomere 4 with anterior angle slightly expanded, with large, thick, strongly curved anterolateral hook-like seta; male protarsomere 5 slightly concave ventrally, with anterior band of ca 100 and posterior band ca 40 of relatively long setae (Fig. 39D); median lobe in lateral view evenly broad, with rounded, not curved downwards apex, in ventral view, evenly tapering, with broadly pointed apex; paramere slightly concave on dorsal side and with long, dense, thin setae, situated along dorsal margin; subdistal setae strong and dense, setae in middle part slightly shorter and sparser, proximal setae long but sparser than subdistal ones (Fig. 39 A-C ). Affinities. In the area of its distribution, E. desii co-occurs with numerous species of the ekari -, ullrichi -, broschii -, larsoni-, and danae -groups. From them, the species differs in size, coloration, surface sculpture, simple male antennae, and the shape of the median lobe. Within the casuarina -group, it is very similar in coloration and surface sculpture to the co-occurring E. pusilla sp. n. and E. pseudopusilla sp. n. but can be distinguished from them by larger size and evidently thicker median lobe. Distribution. Papua New Guinea: East Sepik, Simbu, Western Highlands, and Eastern Highlands Provinces (Fig. 50).