Taxonomic and nomenclatural changes in Cassidinae (Coleoptera: Chrysomelidae)
Author
Sekerka, Lukáš
text
Acta Entomologica Musei Nationalis Pragae
2016
2016-07-15
56
1
275
344
journal article
56063
10.5281/zenodo.5305725
31cb15bf-28f5-4b27-86df-6a1b28cbce41
0374-1036
5305725
E24F1028-C6AC-4323-9ED5-C9B7FF3434ACD
Tribe
Ischyrosonychini Chapuis, 1875
Remarks.
This tribe is assemblage of three groups previously considered as distinct tribes: Asterizini,
Ischyrosonychini
and
Physonotini
.
BOROWIEC (1995)
suggested that all three should be united because of lack of synapomorphies based on adult characters and used the name
Ischyrosonychini
as the valid name.
BOROWIEC (1999a)
also considered the three taxa synonymous, however, used
Physonotini
as the valid name because
Ischyrosonychini
were based on a genus that was a junior synonym of
Eurypedus
Gistel, 1834
. However,
Physonotini
Spaeth, 1942
was proposed after 1930 without description or bibliographic reference to such description thus being unavailable (
ICZN 1999
: Article 13.1) and therefore the name was recently changed again to
Ischyrosonychini
(
BOUCHARD et al. 2011
).
Ischyrosonychini
are superficially similar to
Cassidini
and are quite difficult to define and separate from the latter tribe based solely on adult morphology.
ŚWIĘTOJAŃSKA & WINDSOR (2008)
discussed their position based on larval morphology and concluded that
Ischyrosonychini
possessed features characteristic for primitive (generalized) tribes of Cassidinae s. str. while
Cassidini
were considered as derived. The head of adult specimens of
Ischyrosonychini
is more similar to
Goniocheniini
or
Omocerini
as the head is distinctly transverse with small eyes situated in front of the mouthparts, while
Cassidini
have circular head, mostly with large eyes. In addition all
Ischyrosonychini
have simple and quite long tarsal claws and the ultimate tarsomere is somewhat longer than in most
Cassidini
and as a result they are capable of faster movement than most of
Cassidini
. Also the larvae of
Ischyrosonychini
move more rapidly than those of
Cassidini
(L. Sekerka, unpubl. observation).
Two genera,
Orexita
Spaeth, 1911
and
Aidoia
Spaeth, 1952
, currently classified in
Cassidini
, have a strongly transverse head with small eyes and similar structure of tarsi as
Ischyrosonychini
and I transfer them to the latter tribe. The affinity of the two genera to
Ischyrosonychini
as well as affinities of the genera within the tribe should be verified by molecular data. The genera of
Ischyrosonychini
can be recognized using the key below.