Revisions of Holocnemus and Crossopriza: the spotted-leg clade of Smeringopinae (Araneae, Pholcidae) Author Huber, Bernhard A. 33607F65-19BF-4DC9-94FD-4BB88CED455F Zoological Research Museum Alexander Koenig, Bonn, Germany. b.huber@leibniz-zfmk.de text European Journal of Taxonomy 2022 2022-02-22 795 1 1 241 http://dx.doi.org/10.5852/ejt.2022.795.1663 journal article 10.5852/ejt.2022.795.1663 2118-9773 6299150 7394D45E-46E1-453C-BF7E-1FE1B2CEBB0A Crossopriza soudanensis Millot, 1941 Figs 352 , 354–372 Crossopriza soudanensis Millot, 1941: 28 , fig. 11a–l. Diagnosis Distinguished from known congeners by details of male palp ( Figs 357–361 ; procursus tip with fringed transparent process; shape of distal bulbal sclerite); from the only geographically close congener ( C. illizi sp. nov. ) also by male chelicerae without modified hairs on frontal face ( Fig. 362 ); from many congeners also by female genitalia ( Figs 364–372 ; epigynum with elongate pockets; roundish pore plates far apart; similar to C. illizi ). Type material Lectotype (examined; designated herein) BURKINA FASO • ♂ ; Ouagadougou ; 12.36° N , 1.52° W ; Sep. 1937 ; J. Millot leg.; MNHN Ar 10520. Paralectotypes (examined) BURKINA FASO • 1 ♀ ; same collection data as for lectotype; MNHN Ar 10520 . MALI • 3 ♀♀ , 2 juvs; Bamako ; 12.63° N , 7.99° W ; Oct. 1937 ; J. Millot leg.; MNHN Ar 10512 • 1 ♀ ; Sangha [Sanga]; 14.46° N , 3.30° W ; Oct. 1937 ; J. Millot leg.; MNHN Ar 10518 . Other material examined None. Redescription Male ( lectotype ) MEASUREMENTS. Total length 3.5, carapace width 1.45. Distance PME–PME 80 µm; diameter PME 100 × 140 µm; distance PME–ALE 50 µm; diameter AME 95 µm; distance AME–AME 20 µm. Leg 1: 42.0 (12.0 + 0.7 + 11.1 + 15.9 + 2.3), tibia 2: 7.5, tibia 3: 5.5, tibia 4: 6.1; tibia 1 L/d: 72; femora 1–4 diameters: 0.21, 0.18, 0.17, 0.18. COLOR (in ethanol). Entire specimen very pale, bleached; original coloration ( Millot 1941 ) apparently similar to other epigean congeners. BODY. Habitus similar to C. sahtan sp. nov. and C. ghul sp. nov. (cf. Figs 391 , 481 ). Ocular area slightly raised. Deep thoracic pit and pair of furrows diverging from pit toward posterior margin. Clypeus unmodified, only rim more sclerotized than in female. Sternum wider than long (1.05/0.65), unmodified. Abdomen slightly elongated, dorso-posteriorly barely elevated, rounded. CHELICERAE. As in Figs 362–363 , distally with pair of lateral apophyses provided with one large modified cone-shaped hair each; distance between tips of modified hairs: 390 µm; lateral stridulatory ridges fine but visible in dissecting microscope. PALPS. As in Figs 354–356 ; coxa with strong retrolateral hump; trochanter barely modified; femur small (relative to tibia), distally on ventral side widened, proximally with prolateral stridulatory pick, without retrolateral transversal line, without retrolateral proximal process; femur-patella joints slightly shifted toward prolateral side (arrows in Fig. 354 ); tibia-tarsus joints shifted toward retrolateral side (arrows in Fig. 356 ); tarsus without macrotrichia; procursus ( Figs 357–358 ) short and straight, proximally on prolateral side with hump set with numerous hairs, dorsal hairs straight or curved downwards, procursus tip with ventral sclerite with short dorsal branch, with distinctive process ending in fringed transparent membrane; genital bulb ( Figs 359–361 ) with simple basal sclerite connected to distal (main) sclerite, connecting sclerite with ventral projection, sperm duct opening not seen; distal sclerite with retrolateral ridge, with two small prolateral apophyses and distal rim widely curved towards prolateral. LEGS. Femur 1 with single row of ~26 ventral spines; without curved hairs; few vertical hairs; retrolateral trichobothrium of tibia 1 at 3%; prolateral trichobothrium absent on tibia 1, present on other leg tibiae; tarsal pseudosegments not seen. Female In general similar to male but without spines on legs, apparently without stridulatory files on chelicerae, and with stridulatory organ consisting of pair of small whitish processes posteriorly on carapace and pair of small light brown plates anteriorly on abdomen. Tibia 1 in female from Ouagadougou: 9.1. Epigynum as in Fig. 366 , main epigynal plate oval to semicircular, with pair of distinct elongate pockets (distance between pockets ~310 µm); posterior plate more strongly developed laterally than medially. The only female specimen from Ouagadougou was not cleared. Females from Mali possibly not conspecific with lectotype : very similar epigyna but pockets in female from Sangha slightly closer together ( Fig. 367 ; distance ~250 µm), in females from Bamako considerably farther apart ( Fig. 370 ; distance ~450 µm). Internal genitalia ( Figs 371–372 ; specimen from Bamako) with almost round pore plates, simple dorsal and ventral arcs, ventral arc medially barely modified. Tibia 1 in 2 females from Bamako: 7.8, 8.7 (missing in other females from Mali ). Distribution Known from three localities in Burkina Faso and southern Mali ( Fig. 352 ).