A morphological phylogenetic analysis and generic revision of Australian Helicarionidae (Gastropoda: Pulmonata: Stylommatophora), and an assessment of the relationships of the family 2462
Author
Hyman, Isabel T.
Author
Ponder, Winston F.
text
Zootaxa
2010
2010-05-14
2462
1
1
148
https://biotaxa.org/Zootaxa/article/view/zootaxa.2462.1.1
journal article
10.11646/zootaxa.2462.1.1
11755334
5312710
Caldwellia angularis
(
Férussac, 1821
)
Figures 22K
,
29A
,
31D–F
Material examined.
Mauritius
:
AM
C205299
(
one specimen
dissected, radula examined), E slopes of
Mt. Bambous
,
W of Grand Sable
,
E Mauritius
, on ‘Liane Cerf’, native forest,
400 m
alt.,
Dec. 2000
,
O.L. Griffiths
,
V
.
Florens
.
FIGURE 31.
Radulae.
A–C.
Caldwellia imperfecta
, AM C
205300.
D–F.
Caldwellia angularis
, AM C
205299.
G–I.
Ctenophila caldwelli
, AM C
205301.
J–L.
Erepta odontina
, AM C
205302.
M–O.
Kalidos balstoni
, AM C
205303.
P–R.
Madagascan species MA1, AM C205304. Scale bars 10 mm (A–C), 20 mm (D–L, P–R), 50 mm (M–O).
Description.
External morphology
: Shell 4.1 whorls, orange-brown; whorl profile rounded above and below angular periphery; otherwise shell shape and sculpture as for genus. Animal cream. Mantle laps and lobes as for genus. Caudal apparatus as for genus; caudal horn large; caudal foss diamond-shaped slit in tail.
Mantle cavity
: As for genus. Mantle with visible minor blood vessels.
Digestive system
: As for genus.
Genital system
(
Figures 22K
,
29A
): As for genus. Free oviduct long, internally with partially fused pustules. Bursa copulatrix internally with longitudinal pilasters. Vagina internally with wavy longitudinal pilasters. Penis short; epiphallus enters penis through simple pore; nine longitudinal penis pilasters present; penial diverticulum absent. Penis retractor muscle attached to epiphallus. Epiphallus much longer than penis, simple internally. Epiphallic retractor caecum absent.
Radula
(
Figure 31D–F
): As for genus. Radular formula estimated to contain at least 400 teeth per half-row and at least 200 rows.
Remarks.
Caldwellia angularis
grouped with
Helicarion mastersi
and
Brevisentis atratus
in the phylogenetic analysis (
Figures 1–3
). However, in the molecular phylogenetic analysis of
Hyman
et al.
(2007)
it usually grouped with a different clade of Australian taxa, made up of three species from Lord Howe Island and
Nitor circumcincta
from southeastern
Australia
.