<document id="93ADCA5A5CA1755168D01DE91AC9A009" ID-CLB-Dataset="30595" ID-DOI="10.11646/zootaxa.4388.4.1" ID-GBIF-Dataset="a884362d-a9f8-4cee-8cd4-dcf59a73ede2" ID-ISSN="1175-5326" ID-Zenodo-Dep="1189072" ID-ZooBank="7A57C29B-6B16-445E-A8D7-750C736E6B42" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1520318467094" checkinUser="plazi" docAuthor="Cupello, Mario" docDate="2018" docId="5C282C04FFCFFFEDFF2D50B01C82FA83" docLanguage="en" docName="zootaxa.4388.4.1.pdf" docOrigin="Zootaxa 4388 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Canthidium (Canthidium) vespertinum Howden &amp; Young 1981" docType="treatment" docVersion="10" lastPageNumber="465" masterDocId="A011547CFFC1FFE3FFBA541B1E1AFFFB" masterDocTitle="On the types species of the New World dung beetle genus Canthidium Erichson, 1847 (Coleoptera: Scarabaeidae: Scarabaeinae), with an annotated checklist of species" masterLastPageNumber="486" masterPageNumber="451" pageNumber="465" updateTime="1698528354521" updateUser="ExternalLinkService">
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<mods:title id="EA7D4482CACD3A68933E03405AB1C98E">On the types species of the New World dung beetle genus Canthidium Erichson, 1847 (Coleoptera: Scarabaeidae: Scarabaeinae), with an annotated checklist of species</mods:title>
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<taxonomicName id="1381E691FFCFFFEDFF2D50B01D66FB3E" ID-CoL="QNBB" authority="Howden &amp; Young, 1981" authorityName="Howden &amp; Young" authorityYear="1981" box="[151,892,1195,1221]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="vespertinum" subGenus="Canthidium">
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<emphasis id="E6F54100FFCFFFEDFF2D50B01F38FB3F" bold="true" box="[151,290,1195,1220]" italics="true" pageId="14" pageNumber="465">Canthidium</emphasis>
(
<emphasis id="E6F54100FFCFFFEDFE8850B01FA7FB3F" bold="true" box="[306,445,1195,1220]" italics="true" pageId="14" pageNumber="465">Canthidium</emphasis>
) 
<emphasis id="E6F54100FFCFFFEDFE7650B01C40FB3F" bold="true" box="[460,602,1195,1220]" italics="true" pageId="14" pageNumber="465">vespertinum</emphasis>
<bibRefCitation id="B010E0E3FFCFFFEDFDDB50B71D66FB3E" box="[609,892,1196,1221]" pageId="14" pageNumber="465" refId="ref27924" refString="Howden, H. F. &amp; Young, O. P. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article">Howden &amp; Young, 1981</bibRefCitation>
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<emphasis id="E6F54100FFCFFFEDFF7D50CB1FC4FB1C" box="[199,478,1232,1256]" italics="true" pageId="14" pageNumber="465">Canthidium vespertinum</emphasis>
<treatmentCitation id="5520BB03FFCFFFEDFE5250D41D28FB13" author="" box="[488,818,1231,1256]" page="86" pageId="14" pageNumber="465">
<bibRefCitation id="B010E0E3FFCFFFEDFE5250D41D12FB13" box="[488,776,1231,1256]" pageId="14" pageNumber="465" refId="ref27924" refString="Howden, H. F. &amp; Young, O. P. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article">Howden &amp; Young, 1981</bibRefCitation>
: 86
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[here transferred to the subgenus 
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<emphasis id="E6F54100FFCFFFEDFB7950CB1B5DFB13" box="[1219,1351,1232,1256]" italics="true" pageId="14" pageNumber="465">Canthidium</emphasis>
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]. 
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locality:
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Parama: 
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: Renascimiento, 
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north of Santa Clara, Hartmann’s Finca, 
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, 
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, 
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(“ 
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, 
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Prov., 
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N Sta. Clara, 
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, 
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, 
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”). 
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material:
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<typeStatus id="0B3A23B0FFCFFFEDFB4C510C1B44FACB" box="[1270,1374,1303,1328]" pageId="14" pageNumber="465" type="holotype">Holotype</typeStatus>
male (CMNC—ex Howden collection); see full list of 
<typeStatus id="0B3A23B0FFCFFFEDFD7D51261D29FAAE" box="[711,819,1341,1365]" pageId="14" pageNumber="465" type="paratype">paratypes</typeStatus>
in 
<treatmentCitation id="5520BB03FFCFFFEDFCE051271AB6FAAE" author="" box="[858,1196,1340,1365]" page="86" pageId="14" pageNumber="465">
<bibRefCitation id="B010E0E3FFCFFFEDFCE051271AB6FAAE" box="[858,1196,1340,1365]" pageId="14" pageNumber="465" refId="ref27924" refString="Howden, H. F. &amp; Young, O. P. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article">Howden &amp; Young (1981: 86)</bibRefCitation>
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. 
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<collectingCountry id="AC96DD82FFCFFFEDFAE651271ED1FA83" name="Costa Rica" pageId="14" pageNumber="465">Costa Rica</collectingCountry>
and 
<collectingRegion id="164553F0FFCFFFEDFEBB517A1F40FA83" box="[257,346,1377,1400]" country="Panama" name="Panama" pageId="14" pageNumber="465">
<collectingCountry id="AC96DD82FFCFFFEDFEBB517A1F40FA83" box="[257,346,1377,1400]" name="Panama" pageId="14" pageNumber="465">Panama</collectingCountry>
</collectingRegion>
(Kohlmann &amp; Solís 2006a).
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