Cave millipedes of the United States. XV. Coloradesmus gen. nov. (Diplopoda, Polydesmida, Macrosternodesmidae), and four new species from caves in Colorado, USA
Author
Shear, William A.
Author
Steinmann, David B.
text
Subterranean Biology
2019
32
15
32
http://dx.doi.org/10.3897/subtbiol.32.38161
journal article
http://dx.doi.org/10.3897/subtbiol.32.38161
1314-2615-32-15
731EF530E43D4FB8AAA39B4A0AD1AE09
Coloradesmus
hopkinsae Shear & Steinmann
sp. nov.
Figs 9-10
,
11-14
,
15
Types
.
Male holotype and male paratypes from Colorado, Garfield Co.: Bair Cave, 9800' asl,
39°38'N
,
107°14'W
, 200 ft from entrance, collected September 1998 and 19 June 1999 by D. Steinmann.
Diagnosis.
Similar in somatic characters to both
C. aquiliensis
and
C. manitou
, but differing from both in the broader, more irregularly shaped prefemoral process of the gonopods, and in having a short endomerite tightly appressed to the prefemoral process.
Etymology.
The species is named in honor of Kay Hopkins of the United States Forest Service for her contributions to cave conservation, cave research and cave preservation.
Description.
Male holotype.
Length about 7.0 mm, greatest width 0.66 mm. With the characters of the genus. Head sparsely setose. Basal joint of mandibles not greatly exceeding width of collum; front margin of collum evenly arcuate, posterior margin shallowly embayed (
Fig. 9
). Setae of collum and metazonites acute throughout (
Figs 9
-
11
), setal rows of metazonites sometimes interrupted, supplemental setae sometimes present (
Fig. 10
). Setal tubercles become more prominent posteriorly (
cf.
Figs 10
,
11
). Telson rounded, with two distinct rows of setae; epiproct blunt (
Fig. 11
). Gonopod socket occupies entire width of prozonite, subcordate, indenting ninth sternite (
Fig. 12
). Gonopods with robust acropodite (a,
Fig. 13
), ending in long, acute distal zone (dz,
Figs 13
,
14
); seminal pore long, with many tiny finger-like cuticular projections (
Fig. 15
); endomerite (e,
Fig. 13
) short, tightly appressed to prefemoral process. Prefemoral process (pfp,
Figs 13
,
14
), large, broad, scoop-shaped, with subapical lateral tooth, tip acute, slightly lobed.
Figures 11-14.
Coloradesmus hopkinae
male.
11
Telson, dorsal view
12
gonopods, ventral view
13
right gonopod, mesal view
14
right gonopod, medioposterior view. See text for explanation of labels.
Figures 15-20.
Coloradesmus
species males.
15
C. hopkinae
, gonopod seminal pore
16-20
C. manitou
16
collum, dorsal view
17
midbody ring, dorsal view
18
telson, dorsal view
19
gonopods, ventral view
20
Gonopods, posterior view. See text for explanation of labels.
Distribution.
COLORADO: Garfield Co.: Groaning Cave,
39°42'N
,
107°10'W
, 9800' (2804 m) asl, 9 October 2008, D. Steinmann, male, females, juveniles.