A revision of Frutillaria Richards and Penola Richards (Diptera: Sphaeroceridae: Archiborborinae) Author Kits, Joel H. Author Marshall, Stephen A. text Zootaxa 2011 2863 1 34 journal article 10.5281/zenodo.203208 85473c5c-8506-4915-9dd9-0317382ba608 1175-5326 203208 Frutillaria calceata Richards ( Figs. 10–12 , 63, 64, 67 , 110 ) Frutillaria calceata Richards, 1961 : 67 Description. Length 1.7–2.5 mm (head + thorax 0.8–1.2 mm) Occiput dark brown, face orange, frons and gena intermediately coloured. Shiny spots on frons lateral to ocellar triangle. Scape and first flagellomere orange, pedicel brown. Clypeus orange, prementum brown. Thorax black, postalar calli reddish. Mesoscutum and scutellum entirely covered with microtomentum. Wing rudiment light brown. Anepisternum mostly shiny, anterior, posterior, and dorsal margin with microtomenum. Katepisternum with a shiny spot behind coxa, not extending toward bristle; posterior part of katepisternum, meron and metapleuron shiny with exception of a stripe of microtomentum between posterior thoracic spiracle and base of hind coxa. Laterotergite mostly shiny. Legs black, trochanters, joints, and tarsi orange, fore coxa dark reddish brown. Fore femur covered with microtomentum. Mid femur with 2 anterior bristles. Male abdomen ( Figs. 10–12 ): Tergites 1–5 covered with microtomentum, shiny along the posterior margins. Sternite 2 shiny, sternites 3–4 mostly shiny with microtomentum medially, sternite 5 extensively covered with microtomentum. Ring-shaped sclerite present. Synsternite 6+7 connected by a narrow band to the accessory sclerite flanking the distiphallus. Surstylus with a long inner arm, subequal to length of basal part; outer corner strongly produced and broadly rounded. Postgonite bilobed, anterior lobe very short and thin, posterior lobe strongly angled anteriorly. Basiphallus with both epiphallus and preepiphallus well-developed. Distiphallus with ventroapical discs very large, extending beyond central projecting sclerite; central projecting sclerite rectangular, extending both dorsally and ventrally to lateral arm; lateral arm slightly longer than central projecting sclerite, pointed at tip; ventral sac strongly sclerotized, projecting perpendicularly. Female abdomen ( Figs. 63, 64, 67 ): Tergites 1–6 covered with microtomentum, shiny along the posterior margins. Sternite 2 shiny, sternites 3–4 mostly shiny with microtomentum medially, sternites 5–6 mostly covered with microtomentum, shiny along posterior margins. Tergite 7 distinctly convex posteriorly, with a medially interrupted band of microtomentum along posterior margin. Sternite 7 somewhat pointed medially, mostly shiny with a medial patch of microtomentum. Tergite 8 divided medially, epiproct triangular with fairly long anterior arms. Cerci without microtrichia dorsally. Sclerites of sternite 8 subrectangular, hypoproct with lateral sclerotized spots and a medial sclerotized bar. Spermathecae with long, mushroom-shaped apical invagination, short basal invagination, both annulated. Type material. Holotype 3: CHILE : [ Los Lagos: ] Los Riscos, 450 m , 14 Sep 1954 , [G. Kuschel] ( MEUC , apparently lost). Paratype : Llanquihue Prov., Frutillar, 41º 12' S , 25 Sep 1954 , G. Kuschel (1 3, BMNH , examined). Other material examined. CHILE : Aisén: Queulat Pass, 40 km N Puerto Cisnes, mossy beech forest, carrion trap, 500 m , 30 Dec 1984 – 28 Jan 1985 , S. & J. Peck (1 Ƥ, DEBU ); Los Lagos: Puaucho (15.1 km W), 40°35'00"S 73°37'42"W , 50 m , Valdivian rainforest remnant in sm. ravine, w/large ferns, FMHD #96-248, 30 Dec 1996, berlese, leaf & log litter, A.F. Newton & M.K. Thayer (2 3, 2 Ƥ, FMNH ); as above but FMHD #96-249, berlese, litter under large ferns, A.F. Newton & M.K. Thayer (1 3, 1 Ƥ, FMNH ); Vicente Pérez Rosales Natl. Pk., N slope Volcán Osorno, road to Ref. La Picada, 41°03'15"S 72°30'11"W , 660 m , Nothofagus dombeyi w/conifers, dense Chusquea bamboo understory, FMHD #2002-082, berlese, leaf & log litter, 16 Dec 2002 , Solodovnikov, Thayer & Newton (4 3, FMNH ); Vicente Pérez Rosales Natl. Pk., N slope Volcán Osorno, road to Ref. La Picada, 41°01'03"S 72°32'54"W , 430 m , Nothofagus dombeyi w/conifers, FMHD #2002-083, berlese, leaf & log litter, Newton, Solodovnikov, & Chani-Posse (2 3, 1 Ƥ, FMNH ); Bahia Mansa, 3 km S Maicolpue, 200 m , 21 Dec 1984 – 3 Feb 1985 , carrion trap, mixed forest, S. & J. Peck (1 3, DEBU ); as above but FMHD #85-933, Berlese, litter, 21 Dec 1984 , S. & J. Peck (3 3, 1 Ƥ, FMNH ); as above but FMHD #85-994, Berlese, litter, 3 Feb 1985 , S. & J. Peck (2 3, FMNH ). Comments. The paratype is in poor condition, and the aedeagal and hypandrial complexes are apparently lost. However, based on the surstylus of this specimen and Richards’ figure of the species, it is clear that the more recently collected material belongs to the same species. There is a female (species undetermined) enclosed in a separate glass tube in the vial containing the male paratype , but it is not mentioned specifically by Richards, and is not a paratype .