Three new minute leaf litter beetles discovered by citizen scientists in Maliau Basin, Malaysian Borneo (Coleoptera: Leiodidae, Chrysomelidae) Author Schilthuizen, Menno Author Seip, Lilian A Author Otani, Sean Author Suhaimi, Jadda Author Njunjic, Iva text Biodiversity Data Journal 2017 5 21947 21947 http://dx.doi.org/10.3897/BDJ.5.e21947 journal article http://dx.doi.org/10.3897/BDJ.5.e21947 1314-2828-5-21947 Colenisia Fauvel, 1903; Type species: Colenisia caledonica Fauvel, 1903 Colenisia Fauvel, 1903; ( Fauvel 1903 , Newton 1998 , Daffner 1991 , Svec 2013 ) Colenisia Colenisia caledonica Fauvel, 1903 Fauvel 1903 Diagnosis Body small (0.8-2.5 mm), oval, glossy, usually uniformly brown. Head broad, occipital crest absent, at least half as wide as the pronotum, with distinct microreticulation. Antennal insertion concealed, antennal groove absent. Antennae 11-segmented, relatively compact, 8th antennomere much smaller than 9th and 10th. Labrum not emarginate; mandible with strongly developed molar surface. Elytra with transverse microreticulation. First abdominal segment without a transverse carina. Hind coxae not separated. Tarsal formula 5-4-4 in both sexes. Aedeagus with free parameres.