Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae) Author PRATHER, AYSHA L. text Zootaxa 2003 2003-08-29 275 1 1 214 https://biotaxa.org/Zootaxa/article/view/zootaxa.275.1.1 journal article 5618 10.11646/zootaxa.275.1.1 9322f2b2-a384-40bf-9d57-d789c76ce5c4 1175­5334 5019235 89BC39FD-ABCC-4634-876D-8C1F4337C325 Phylloicus maculatus (Banks) Figs. 64 , 65 Phylloicus maculatus ( Banks, 1901:369 ) [ Type locality: Mexico , Veracruz , Presidio; MCZ; female; in Heteroplectron ]. — Flint 1967:19 [female, to Phylloicus ]. — Holzenthal 1988:72 [distribution]. Phylloicus maculatus is distinguished by its large size, wing pattern, and by having short, single­lobed lateral coremata, without modifications of tergite IV ( Fig. 64F ). The wing pattern of this species is identical to that of P. elegans and lituratus . The female genitalia are indistinguishable from those of the latter two species. Adult. Forewing length 13.415. 4 mm , n = 31. Head dark brown, setal warts very dark. Maxillary palps dark brown, covered with golden brown setae on basal three segments. Antenna twice forewing length; dark brown, with narrow patches of pale sensilla on anteromesal surface of each flagellomere. Dorsal pterothorax dark brown, mesoscutellum golden brown; ventrolateral thorax golden. Femora golden; tibiae golden; tarsi dark brown. Metathoracic leg of male with posterior fringe of long setae. Tibial spur formula 2,4,4. Forewing flat; dark brown; with two transverse bands; proximal band ivory, reaching posterior wing margin, at least 1/2 width of wing, an inverted V­shape; distal band ivory, beginning at anterior wing margin, at least 1/2 width of wing; with two basal stripes, ivory. Hind wing basal brush present in male and female; female brush short. Male. Preterminalic abdominal terga with anteromesal notch. Corematic structures present. Tergum IV with expanded lateral flanges, lateral coremata; lateral coremata single­lobed and short, cylindrical. Tergum V without sclerotized modifications ( Fig. 64F ). Sternum VII with short, acute anteromesal process. Sternum VIII enclosing base of elongate sternum IX; posteromesal process notched ( Fig. 64A, C ). Tergum IX with paired mesal ridges extending 1/2 length of segment; posterior margin with round narrow mesal projection ( Fig. 64B ); lateral ridge absent; dorsal pleural setae approximately 5, ventral pleural setae approximately 8 ( Fig. 64A ). Preanal appendage longer than tergum X, but less than 11/2 times length, widest apically, setae filamentous, longer than appendage ( Fig. 64A, B ). Tergum X without basal lobes; basodorsal process short and digitate; basolateral processes long, length at least twice diameter; apex, in lateral view, rounded, in dorsal view, entire ( Fig. 64A, B ). Harpago rounded; peglike setae many, apical ( Fig. 64A, C ). Phallic endotheca with paired basolateral lobes, basolateral lobes tapered apically; phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoe­shaped ( Fig. 64D, E ). Female. Preterminal abdominal terga with anteromesal notch. Sternum VII with short pointed anteromesal process. Tergum VIII without posterolateral brush; sternum VIII with shallow posteromesal notch, or posterior margin entire; sternum VIII ( Fig. 65C ). Tergum IX without mesal ridge ( Fig. 65B ). Sternum IX anterior and posterior lobes darkly sclerotized and striate, with patch of lightly sclerotized cuticle lateral to vaginal opening ( Fig. 65A ). Tergum X appendage shorter than mesal lobe, base marked by faint suture line, apex oblique; mesal lobe lightly sclerotized; digitate lateral processes length approximately equal diameter ( Fig. 65B ). Sternum X with patches of short fine setae posterolaterally to anal opening ( Fig. 65A ). Vaginal apparatus anterior and posterior sclerites equal in length; anterior sclerite rounded anteriorly, posterolateral projections absent; posterior sclerite triangular; posterior end of spermatheca a sclerotized ovoid ( Fig. 65A ). Material examined. GUATEMALA : Izabal : Cayuga , 15°32'00”N , 88°42'00"W , 1.ii.1915 , Schaus — 1 female ( NMNH ) ; MEXICO : Veracruz : Presidio , 19°05'00”N , 96°58'00"W , 1.vi. holotype female ( MCZ ) ; Municipio Tilapan , Arroyo Quetzalan. Los Manantiales km 5 Rta. Orizaba­Zongolica , 18°47'28”N , 97°06'05"W , 1160 m , 17.iii.2000 , Bueno , Barba , & Rojas — 1 female ( UMSP ) ; 13.vii.2001 , Bueno & Barba — 12 males , 10 females ( IBUNAM ) ; — 3 males , 3 females ( UMSP ) . Distribution. Guatemala , Mexico