Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae) Author PRATHER, AYSHA L. text Zootaxa 2003 2003-08-29 275 1 1 214 https://biotaxa.org/Zootaxa/article/view/zootaxa.275.1.1 journal article 5618 10.11646/zootaxa.275.1.1 9322f2b2-a384-40bf-9d57-d789c76ce5c4 1175­5334 5019235 89BC39FD-ABCC-4634-876D-8C1F4337C325 Phylloicus elegans Hogue and Denning Figs. 43 , 44 , 109 Phylloicus elegans Hogue and Denning, 1983 in Denning et al. 1983:184 [ Type locality: Panama , Canal Zone, Barro Colorado Island; WSU; male]. — Flint 1991:98 [male, distribution]. The wing pattern of P. elegans ( Fig. 109 ) is identical to that of P. lituratus . However, the male abdomen of P. elegans is very distinctive: tergum IV has distinctively shaped posteriorly projecting lateral sclerites and multilobed lateral coremata ( Fig. 43F ). The females of the two species are indistinguishable, and as several series from different localities contain males of both P. elegans and P. lituratus , all determinations of females are tentative. Hybrids may occur; this is discussed under P. lituratus . Adult. Forewing length 10.4­12.1 mm, n = 106. Head chestnut brown. Maxillary palps dark brown. Antenna twice forewing length; chestnut brown, with narrow patches of pale sensilla on anteromesal surface of each flagellomere. Dorsal pterothorax chestnut brown; ventrolateral thorax golden. Femora golden; foretibiae dark brown; mesotibiae dark brown; metatibiae dark brown; foretarsi dark brown; mesotarsi white proximally, dark distally; metatarsi dark brown. Metathoracic leg of male with posterior fringe of long setae, setae dark. Tibial spur formula 2,4,4. Forewing flat; dark brown; with two transverse bands; proximal band ivory, reaching posterior wing margin, at least 1/2 width of wing, an inverted V­shape; distal band ivory, beginning at anterior wing margin, at least 1/2 width of wing; with two basal stripes, ivory ( Fig. 109 ). Hind wing basal brush present in male and female; male brush light brown; female brush dark brown. Male. Preterminalic abdominal terga with anteromesal notch. Corematic structures present. Tergum IV with paired posterior processes and paired lateral sclerites, lateral coremata; posterior process truncate; lateral sclerite spatulate, directed laterally; lateral coremata with basal globose lobes. Tergum V without sclerotized modifications ( Fig. 43F ). Sternum VII with short, acute anteromesal process. Sternum VIII enclosing base of elongate sternum IX; posteromesal process notched, notch deep and round ( Fig. 43A, C ). Tergum IX with mesal ridge extending full length of segment; posterior margin with round narrow mesal projection ( Fig. 43B ); lateral ridge present; dorsal pleural setae approximately 4, ventral pleural setae approximately 2 ( Fig. 43A ). Preanal appendage at least 11/2 times length of tergum X, widest apically, setae filamentous, longer than appendage ( Fig. 43A, B ). Tergum X without basal lobes; basodorsal process short and digitate; basolateral processes short, length less than or equal to diameter; apex, in lateral view, truncate, in dorsal view, entire or notched, notch shallow; with paired dorsolateral ridges ( Fig. 43A, B ). Harpago rounded; peglike setae many, apical ( Fig. 43A, C ). Phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoe­shaped ( Fig. 43D, E ). Female. Preterminal abdominal terga with anteromesal notch. Sternum VII with short pointed anteromesal process. Tergum VIII without posterolateral brush; sternum VIII with shallow posteromesal notch, or posterior margin entire; sternum VIII ( Fig. 44C ). Tergum IX without mesal ridge ( Fig. 44B ). Sternum IX anterior and posterior lobes darkly sclerotized and striate, with patch of lightly sclerotized cuticle lateral to vaginal opening ( Fig. 44A ). Tergum X appendage shorter than mesal lobe, base indistinct, apex oblique; mesal lobe lightly sclerotized; digitate lateral processes absent ( Fig. 44B ). Sternum X with patches of short fine setae posterolaterally to anal opening ( Fig. 44A ). Vaginal apparatus anterior and posterior sclerites equal in length; anterior sclerite truncate anteriorly, posterolateral projections absent; posterior sclerite triangular; posterior end of spermatheca a sclerotized cone ( Fig. 44A ). Material examined. COLOMBIA : Antioquia : Quebrada Honda, 12 km SW Fredonia, 1450 m , 22.ii.1983 , Flint — 3 males , 1 female ( NMNH ) ; Quebrada La Jimenez , Mun. Sopetran , trap C , 780 m , 22.v.1983 , Matthias — 1 male ( NMNH ) ; COSTA RICA : Alajuela : 20 km S Upala , 15.vii.1990 , Parker — 2 males ( EMUS ) ; 6.i.1991 , Parker — 1 male ( EMUS ) ; 30.v.1991 , Parker — 1 male ( EMUS ) ; 1­15.vii.1991 , Parker — 1 male ( EMUS ) ; 1­10.viii.1991 , Parker — 1 male ( EMUS ) ; 21­31.viii.1991 , Parker — 1 male , 1 female ( EMUS ) ; unnamed river, Cerro Campana ca. 6 km (air) NW Dos Rios , 10°54'00”N , 85°24'00"W , 640 m , 22­23.vii.1987 , Holzenthal , Morse , & Clausen — 1 male ( UMSP ) ; Guanacaste : Maritza , Quebrada Marilin , 10°57'04”N , 85°29'24"W , 600 m , 12.i.1992 , Sweeney , MacLeod , & Villalobos — 3 males ( NMNH ) ; 9.xi.1992 , de la Rosa — 1 male ( UMSP ) ; Río Los Ahogados , Río Los Ahogados , 11.3 km ENE Quebrada Grande , 10°51'54”N , 85°25'23"W , 470 m , 7.iii.1986 , Holzenthal & Fasth — 1 male ( UMSP ) ; Parque Nacional Guanacaste , Río Orosí , Estación Pitilla , 10°59'28”N , 85°25'41"W , 700 m , 22­25.v.1990 , Holzenthal & Blahnik — 1 male ( UMSP ) ; Límon : Río Uatsi , ca. 8 km (air) W Bribri , 09°37'12”N , 82°54'00"W , 60 m , 25.v.1987 , Holzenthal , Hamilton , & Heyn — 1 male ( UMSP ) ; Puntarenas : Río Singrí , ca 2 km (air) S Finca Helechales , 09°03'25”N , 83°04'55"W , 720 m , 21.ii.1986 , Holzenthal , Morse , & Fasth — 1 male ( UMSP ) ; San José : Escazu , 11­18.iv.1988 , Parker — 2 males ( EMUS ) ; 15.iv.1988 , Parker — 2 males ( EMUS ) ; 15­21.v.1988 , Parker — 1 male , 1 female ( EMUS ) ; Reserva Biológica Carara , Río Carara , Carara , 09°46'41”N , 84°31'52"W , 200 m , 14.iii.1991 , Holzenthal , Muñoz , & Huisman — 1 male ( UMSP ) ; Río del Sur , 1.5 km (rd) S Carara , 09°46'08”N , 84°31'52"W , 160 m , 13.iii.1991 , Holzenthal , Muñoz , & Huisman — 1 male ( UMSP ) ; ECUADOR : Pichincha : Río Palenque Biological Station , Río Palenque , Santo Domingo ( 47 km ) , 229 m , 29.vii.1976 , Cohen — 1 male ( NMNH ) ; NICARAGUA : Chontales : Rt. 7, km 206 E of Villa Somosa , 29.vii.1967 , Flint & Ortiz — 1 male ( NMNH ) ; Jinotega : Cerro Mazú , 14°33'00”N , 85°07'00"W , 220 m , 7­10.ix.1997 , Maes & Hernández — 2 males , 1 female ( UMSP ) ; Zelaya : Cerro Saslaya , 13°44'00”N , 85°01'00"W , 700 m , 1.iv.1996 , Maes & Hernández — 1 male , 1 female ( NMNH ) ; PANAMA : Darien : Río Tacarcuna , 579 m , 11.vii.1963 , Fairchild — 1 male ( MCZ ) ; Panama : Canal Zone, Barro Colorado Island , 28­ 30.iv.1964 , S & W Duckworth — 1 male ( NMNH ) ; 10­17.v.1964 , S & W Duckworth — 1 male ( NMNH ) ; 21.ii.1967 , Akre — 12 male , 21 female paratypes ( CAS ) ; 22­24.ii.1967 , Akre — holotype male ( WSU ) ; 12.iii.1967 , Akre — allotype female ( WSU ) ; 18.iii.1967 , Akre — 10 male , 14 female paratypes ( CAS ) ; 24­25.ii.1969 , Akre — 8 male , 9 female paratypes ( CAS ) ; 12.iii.1969 , Akre — 1 male , 2 female paratypes ( CAS ) ; 8.v.1977 , Silberglied & Aiello — 1 male ( NMNH ) ; 25.vi.1978 , Silberglied & Aiello — 1 male ( NMNH ) ; Snyder­Molino trail, marker 3 , 28.ix.1988 ­ 3.i.1989 , Wolda 7 males ( NMNH ) ; 13.xii.1989 ­ 13.ii.1990 , Wolda — 6 males ( UMSP ) ; 12.xii.1990 ­ 5.ii.1991 , Wolda — 5 males ( NMNH ) ; Windowpane trap­4A , 31.i.­5.ii.1986 , Wolda — 1 male ( NMNH ) . Distribution. Colombia , Costa Rica , Ecuador , Nicaragua , Panama .